Wernham .— The Morphology of 
tapering to a point. From the midst of the leaves arises the long 
peduncle, bearing terminally a small ovoid strobilus with closely over¬ 
lapping bracts, spirally arranged. 
Internal Anatomy. 
Hie. 
The internal anatomy of Phylloglossinn has been exhaustively de¬ 
scribed by Bertrand ( 5 ); in the present paper only the broader features 
will be considere . 
The succeeding description will follow the microtomed sections from 
below upwards ; the first structure which presents itself is :— 
i. The Young Tuber. At the extreme tip this consists solely of an 
aggregate of parenchymatous cells, with well-marked nuclei ; but at a very 
short distance above, this begins to be clearly differentiated from the 
peripheral layer, which is of regular, more or 
less radially elongated, empty cells. This 
peripheral layer persists throughout the entire 
length of the tuber ; followed upwards, it pre¬ 
sents a more or less strong thickening of the 
exterior cell-walls. Sections of the older 
tuber, at a higher level, also reveal a well- 
marked thickening on the radial walls of the 
peripheral cells. 
Ascending, the sections show a gradual 
aggregation of the nucleated cells towards 
the centre, these being surrounded by a broad 
zone of homogeneous parenchyma, in which 
intercellular spaces are noticeably absent. The latter zone is doubtless 
destined to form the food-storage tissue, the nucleated cells being prob¬ 
ably merismatic (Fig. 2). 
Proceeding upwards, the central merismatic zone becomes smaller and 
smaller, ultimately breaking loose from the surrounding parenchymatous 
zone, and coming to lie in a cavity. 
In the meantime the radial walls of the peripheral cells have acquired 
a well-pronounced thickening, particularly in the centre, so that they present 
a resemblance to the appearance often associated with an endodermis. 
The next twelve sections in ascending order show simply a central 
cavity—indicating the existence of a rather long central channel (Fig. 4) 
in the ‘ stalk ’ of the tuber. This is borne out by the longitudinal sections, 
as shown in the diagram (Fig. 3). 
One or two of the nucleated cells persist for a time around the edge 
of the central cavity as the transverse sections are followed upwards, so 
that the meristem tapers to a fine point (Figs. 3, 4); the segmentation 
Fig. 2. Diagram of trans¬ 
verse section of young tuber; pe, 
peripheral layer; pa , storage paren¬ 
chyma ; me , merismatic tissue. 
