Phylloglossu m Drum mondii. 
34i 
The position of the phloem in this stele would be of the most significant 
interest, but the closest examination has failed to reveal its presence in the 
stem. 
5. The Peduncle of the Strobilus. This follows in the transverse sections 
in a continuous ascent from the stem region. The xylem of the latter 
breaks up, very soon after entering the peduncle, into isolated strands, very 
irregularly disposed ; not in a circle, as has apparently been the general 
impression hitherto (Fig. 8). 
Each strand encloses a protoxylem-cavity and is thus clearly mesarch 
in development. One or 
two small wood-elements 
cling to the edge of the / 
cavity ; and here the re¬ 
semblance to the carinal 
canals of Equisetum is 
much more striking than 
in the stem-stele. 
6. The Strobilus re¬ 
gion. As we approach the 
base of the strobilus, one 
or two of the strands move 
outwards to the periphery 
of the section ; these, we 
shall see, go to supply 
sporophylls. The first in¬ 
dication of the latter in the 
sections is the appearance 
of a tangentially elongated 
mass of parenchyma, en¬ 
tirely isolated from the 
section of the main axis, 
and destitute of vascular 
tissue. This is obviously the down-pointing free ‘ dorsal lobe ’ of the 
lowest sporophyll 1 (dl y Fig. 8). Opposite this, in the axis section and 
near its periphery, is the xylem-strand (/) which is going to supply the 
sporophyll. 
Fig. 8 shows the general appearance seen in a transverse section 
taken about midway across the strobilus. Dorsal lobes (dl), sporophylls (. sp ), 
and sporophyll traces are seen in section at different levels. The sporophylls, 
like the leaves, are seen to possess a small central vascular strand, the latter 
originating from the isolated bundles of the axis ; one is shown, in partial 
longitudinal section, in the act of passing out into its sporophyll. 
1 Sykes ( 11 ). 
Fig, 8 . Transverse section of strobilus. The walls of 
seven sporangia are seen in section, sw. The section passes 
through the base of one sporophyll, before it has become free 
of the main axis. Xylem black, parenchyma shaded. 
