350 Cutting .— On Androgynous Receptacles in Marchcintia . 
There were present two kinds of archegoniophore, the disc-type with 
short processes, and the ‘ polymorpha ’ type with long processes ; and two 
kinds of antheridiophore—one with only a slightly crenate edge and equal 
lobes, and one that was asymmetric and in which the lobes were free for 
the greater part of their length. In the last mentioned the number of 
lobes varied greatly. 
A microscopic examination of the thallus and gametophores revealed 
the fact that at least three different types of pore were present in the 
material, and this fact, as well as the presence of different types of gameto¬ 
phores, makes it more than probable that two or more species had somehow 
got mixed. 
I intended to try to get a further supply from Chelsea in the hope 
that more androgynous receptacles might be found, and that experiments 
might be tried in order to find the factors that regulate the appearance 
of this condition in this species, and to find out whether it can be inherited. 
As both Schiffner ( 17 ) and Stephani ( 18 ) regard Marchantia palmata 
and Marchantia emarginata as synonyms, and as two very characteristic 
species under these names are grown at Chelsea, both of these were 
collected during this year: neither of them, however, showed the androgy¬ 
nous condition. The Marchantia palmata formed archegoniophores like 
those with the long processes found in the Avery Hill material, but no 
disc-shaped ones. The M. emarginata formed very few gametophores 
during the year and none that I had were adult, but Mr. Hales, the 
Curator of the Gardens, informs me that the adult shape is in the form 
of a disc similar to that of the androgynous receptacles described above. 
Mr. Gepp, of the Natural History Museum, has very kindly shown me 
the Herbarium specimens of M. palmata and M. emarginata. They seem 
very similar and both are evidently very variable, but neither of them 
resembles either the Chelsea specimens or my own: nor does Stephani’s 
description of M. palmata (syn. M. emarginata) agree with either the 
Chelsea specimens or my material. The latter, unfortunately, is in too 
fragmentary a condition for the separation of the constituent species and 
the determination of the specific name of the androgynous specimens. 
As I have failed to find any androgynous gametophores on the Chelsea 
specimens, and as the androgynous material at my disposal was insufficient 
for the determination of the species, it was determined to publish the 
results that have so far been obtained. 
The disciform receptacle has a wide, slightly concave upper surface, 
and its margin is produced into 6-12 short, blunt protuberances which at 
times show indications of a slight apical depression (Figs, i, 2, and 3). 
Between two of the protuberances on each receptacle is a very deep cut 
reaching in some cases almost to the centre of the disc-like upper surface. 
This cut indicates the first dichotomy of the shoot which gave rise to the 
