352 Cutting .— On Androgynous Receptacles in Marchantia. 
has been seen. In the Marchantia here described we have, therefore, 
a male outgrowth from one or more branches that have not only been 
formed in the manner characteristic of female branches, but which also 
bear archegonia. The outgrowth, as has been mentioned above, may form 
only one branch (Fig. i), or it may divide again to form an asymmetric 
branch-system ; as many as three branches have been seen in one such 
outgrowth. 
The structure of the antheridia and of the branch which bears them 
is perfectly normal, with the usual amphigastria and rhizoids. One very 
remarkable feature, however, was noticed. Although the gametophores 
Fig. 4. Longitudinal section through an androgynous receptacle, showing hermaphrodite branch on 
right side. ? = archegonia, <J = antheridia, j = stalk, and i= involucre, x 15. Semidiagrammatic. 
were provided with a long stalk and gave other evidence of being fairly 
adult, the majority of the antheridia had not yet discharged their contents. 
In the older parts of the male outgrowths empty antheridia were found, 
so that the fact that the majority were full evidently was not caused by 
their not having the power of opening. It seems, on the other hand, 
to suggest that the male outgrowth was formed secondarily as a kind 
of proliferation, and is not a mere replacement of the normally female 
branch. This point will be discussed later. 
Only about half of the disc-shaped gametophores bore male out¬ 
growths, but the latter easily fall off, and it is certain that many have lost 
them. It is probable, however, that some few of them were purely female, 
as no sign of a broken surface was seen on them. 
