Gnomonict ery thro stoma, Pers. 603 
association with ‘ coils ’. It is suggested that the trichogynes, though 
originally the receptive organs, now perform a different function, possibly 
a respiratory one. 
4. The ‘ coils’ are the first beginnings of perithecial development. In 
the centre of each ‘ coil 5 one or more slightly differentiated hyphae are 
found. The latter are considered to be of the nature of ascogonia. No 
clear connexion can be traced between the trichogynes and the ascogonia. 
5. The stages in the development of the perithecium from the ‘ coil ’ 
are described. During these processes it is no longer possible to trace the 
ascogonia. The ascogenous cells appear to arise by differentiation from 
ordinary cells. 
6. The only nuclear fusion that has been observed takes place in the 
young ascus. 
7. The nuclear divisions in the ascus are described. Only a single 
process of reduction has been seen. This occurs in the first division. It is 
not possible to decide whether this division is heterotype or brachymeiotic, 
though the analogies seem to be with the former rather than the latter. 
8. Thus Gnomonia erythrostoma would appear to be an Ascomycete in 
which only a single nuclear fusion and a single reduction occur normally 
in the life-cycle. 
Botany School, Cambridge, 
Jtme, 1910 . 
List of Papers. 
а. Baur, E. (’98) : Zur Frage nach der Sexualitat der Collemaceen. Ber. d. Deut. Bot. 
Gesell., xvi. 
1 . Blackman, V. H. (’04) : On the Fertilisation, Alternation of Generations, and General Cytology 
of the Uredineae. Ann. Bot., xviii. 
2. Blackman, V. H., and Fraser, H. C. I. (’05): Fertilisation in Sphaerotheca. Ann. 
Bot., xviii. 
3 . --- (’06) : On the Sexuality and Development of 
the Ascocarp in Humaria granulata. Proc. Roy. Soc., London, B. lxxvii. 
4. Brefeld, O. (’72 et seq.) : Untersuchungen aus dem Gesammtgebiete der Mykologie. Leipzig. 
5. Christman, A. H. (’05) : Sexual Reproduction in the Rusts. Bot Gaz., xxxix. 
б . Claussen, P. (’07) : Zur Kenntniss der Kernverhaltnisse von Pyronema conflnens. Ber. d. 
Deut. Bot. Gesell., xxv. 
7 . Cutting, E. M. (’09) : The Sexuality and Development of the Ascocarp of Ascophanus carneus. 
Ann. Bot., xxiii. 
8 . Dale, E. (’03) : Observations on the Gymnoascaceae. Ann. Bot., xvii. 
9 . - (’09) : On the Morphology and Cytology of Aspergillus repens. Ann. Myc., vii. 
10 Dangeard, P. (’94-95) : La Reproduction sexuelle des Ascomycetes. Le Botaniste, iv. 
11 . - (’04) : Recherches sur le developpement du Pdrithece ches les Ascomycetes. 
Le Botaniste, ix. 
