732 Dig by .— The Somatic , Premeiotic , and 
linin becomes finer and more granular, it is increasingly confined to 
the periphery. Hence, in order to ascertain the nature of the linin 
network, it is necessary to examine such sections of nuclei which are so 
cut as to show either the ‘ floor ’ or the ‘ roof giving a broad view of 
the concave or convex surface of the network (Fig. 7). In sections through 
the centre of a nucleus the threadwork is shown squeezed against the 
nuclear wall. In such a view there must of necessity appear to be an 
abundance of parallelism in the threads as the sides of the meshes are seen 
in perspective. This is an easy source of error. 
£ Resting ’ stage. 
The chromatic linin framework continues to break up until a rough mesh- 
work results. The finely drawn out linin threads now connect small rounded 
linin granules (Figs. 6 B and 7). The meshes of the network in some places 
may lie in strands, in others they appear stretched like a net. This is 
the nearest approach to a £ resting ’ nucleus. There is no real rest in the 
quickly dividing cells of a young root, or of the archesporium. These so- 
called £ resting ’ stages are not common in the roots and are still more rare 
in the archesporium. Strasburger ( 30 ) (1904) had difficulty in finding 
£ resting ’ nuclei in Galtonia. Possibly in some nuclei this stage may be 
omitted and they may pass on directly into the prophase. In the non¬ 
dividing portions of the tissues, ‘resting’ nuclei with a far finer, almost 
cloudy looking, reticulum can be constantly found. 
It is necessary here to make a slight digression and to mention curious 
crystalline looking bodies (PI. LX, Fig. 28) which are often present in the 
nuclei of the two or three outer rows of cells of the root, and especially in 
the outermost row. They are apparently absent from the more quickly 
dividing cells of the centrally placed tissue. These ‘structures’ seem to be 
confined almost entirely to those nuclei which are £ in rest \ Leitgeb ( 15 ) 
found crystals in all parts of Galtonia candicans except in the underground 
portions of the roots and the bulbs. He described a vacuole round the 
crystals, and this vacuole might increase in size to so great an extent as to 
burst the nucleus, and the crystals were then projected into the cell lumen. 
Fie ascertained that the crystals gave a proteid reaction. Zimmermann ( 36 ) 
has described the presence of crystals in the roots, epidermis, and endo¬ 
sperm, &c., of Ferns and Phanerogams. More recently Walker and Tozer ( 33 ) 
have figured nucleolar budding in the roots of Beans, and have shown that 
the buds pass out into the cytoplasm. In Galtonia these £ structures ’ 
apparently originate from the nucleolus. There may be one, or more, 
present in a nucleus. They are often crystalline in appearance and are of 
various shapes and sizes (Fig. 28). Some are like chips, some like blocks, 
some are oval. Sometimes they are split so as to appear double (Fig. 28 e) 
and then closely resemble a small pair of chromosomes. In every case they 
