Meiotic Nuclear Divisions of Galtonia candicans . 737 
nexions join the various fragments (Figs. 30 and 31). One or, sometimes, 
two nucleoli are formed. At first the nucleolus takes a chromatin stain, 
but it quickly changes this staining reaction and becomes a typical 
eu-nucleolus of a most colourless nature. 
Early Prophase of First Meiotic Division. 
Rapidly the cells of the anther enlarge, and the nuclei become more 
widely separated, and at the same time increase in size. A still finer 
fragmentation of the linin proceeds, resulting in the production of 
variously sized, rounded, and angled particles (Figs. 31 and 33). Many of 
these lie parallel to one another. A very common form is that of parallel 
rows of paired bead-like portions (Figs. 33, 36, and 37). These beads may 
be in single pairs, or several may be strung together by delicate threads. 
Each bead is convex on its outer, and straight on its inner, side. In the 
same nucleus in which some of the linin has thus finely fragmented, there 
may still be found large alveolized bands of chromosomes consisting of a 
broad strip of linin showing a clear centre (Figs. 34 and 35). As in the 
somatic divisions the chromatin appears to be diffused throughout the 
linin. There is a uniform staining capacity. 
Meanwhile the nuclear framework tends to withdraw to the periphery, 
leaving the nucleolus in a more or less clear space in the centre (Fig. 36). 
Owing to the increase in size of the nucleus and to its peripherally arranged 
framework, the nucleus now appears to be emptier than at an earlier stage 
of prophase. Especially in a thin and superficial section (Fig. 35) of the 
framework, striking examples of vacuolization in the linin fragments, and 
of parallelism in the threads, are to be seen. As in the somatic prophases 
a net-like arrangement of fine connexions joins these fragments to one 
another. There is no limit to the variety of forms which the linin 
portions may adopt, but throughout there is a general impression of 
parallelism in the linin threads, or portions of threads, and of vacuolization 
in the less segmented chromosome strands. These strands which show 
vacuolization are portions of chromosomes which have retained their 
entirety to a comparatively greater extent, and whose sides have not yet 
separated. 
Thus except for the size of the nuclei the somatic and the early heterotype 
prophases closely resemble one another, and, as will be subsequently shown, 
the homotype prophases are also similar. The parallel portions in both 
represent longitudinal halves of somatic chromosomes , and are probably the 
sister halves of the same chromosome, which are now severally coming 
together and condensing to form the somatic or univalent chromosomes. 
From here onwards differences in the two types of division occur. In 
the somatic divisions the parallel paired portions of the chromosomes 
become more and more condensed to form each single univalent chromo- 
