Meiotic Nuclear Divisions of Galtonia candicans. 745 
(Fig. 64), the chromosomes contract to their extreme limit, move to the 
periphery of the nucleus, and enter the well-known diakinesis stage. At 
the same time radiations appear in the cytoplasm (Fig. 64). As already 
described by Miyake ( 21 ) the spindle has a multipolar origin and gradually 
becomes bipolar. 
This description of the origin of the univalent chromosomes of the first 
meiotic division agrees fundamentally with that of Strasburger in 1904 ( 30 ) 
and of Mottier ( 22 ). In 1905 Strasburger ( 31 ) and Miyake ( 21 ) both con¬ 
cluded that the spireme as it comes out of synapsis is bivalent, and that the 
two univalent chromosomes are formed by a separation of the two longitu¬ 
dinal halves of the spireme. In this investigation it has been concluded 
that the spireme is univalent, and that the univalent strands only join 
together secondarily to form the bivalent lengths which finally split apart 
into the univalent chromosomes. 
Spindle Figures. 
The fibres push their way into the nucleus, and the chromosomes 
collect on the spindle (Fig. 65). The entire univalent chromosomes of each 
pair move off one to either pole (Fig. 67). A polar view of the equatorial 
plate shows the great variety in size of the chromosomes ; the two small ones 
usually take up a central position (Fig. 66). As the chromosomes approach 
the poles the longitudinal fission is once more to be seen (PI. LXIII, Fig. 68). 
The halves may separate widely so as to form V-shaped figures (Fig. 68). 
The polar view of an aster shows the eight chromosomes longitudinally 
split (Fig. 69). Again the small ones are centrally placed (Fig. 69). 
Anaphase. 
Arrived at the poles, as in the somatic divisions, the chromosomes 
at first form a condensed mass (Fig. 70). Then they separate out, fragment, 
and the fragments become alveolized, showing ‘ windows ’ and parallel 
threads (Fig. 71). One or two nucleoli appear, and a nuclear wall sur¬ 
rounds the daughter nuclei. At the same time a cell-plate is laid down at 
the equator of the spindle (Fig. 72). The refractive granules in the cyto¬ 
plasm round the nuclei and about the spindle are obvious (Fig. 72). Thus 
imperceptibly the anaphase leads into the telophase (Figs. 70, 71, and 72). 
Telophase. 
The telophase of the first meiotic division is precisely like that of the 
somatic divisions (Figs. 71 and 72). There is the same fragmentation, and 
the same alveolization of the fragments resulting in parallel threads (Fig. 73). 
The linin, as a whole, does not break up into such fine portions as in the 
somatic divisions, but remains in rather large fragments, as usual united by 
fine threads. There is no resting stage. Thus the telophase of the hetero- 
type passes into the prophase of the homotype (Fig. 74). 
