Meiotic Nuclear Divisions of Galtonia candicans . 753 
univalent in character, the longitudinal fission in its substance being homo¬ 
logous with that of the presynaptic stages, and consequently with that of 
the somatic prophases. The univalent homologous lengths of spireme may 
have joined end to end, or be partially united during synapsis, but it is not 
until the hollow spireme and second contraction that the pairing and fusion 
of the univalent chromosomes to form the bivalent segments are completed. 
As the bivalent chromosome segments come out of second contraction, they 
split apart into the two univalent chromosomes. At the homotype division 
these univalent chromosomes split longitudinally. 
3. ‘ Crystalline ’ structures are present in the nuclei of the outer two or 
three rows of cells of the roots. 
4. ‘ Chromatic 5 bodies are given off from the nucleus during the pre¬ 
synaptic, synaptic, and hollow-spireme stages. 
In conclusion I wish to express my grateful thanks to Professor J. Bret- 
land Farmer, F.R.S., for the constant and valuable help, advice, and 
criticism that he has given me throughout the course of this work. 
Bibliography. 
1 . Davis, B. M. (’09) : Pollen Development of Oenothera grandijiora. Ann. of Bot., xxiii, No. 92, 
Oct., pp. 551-71. 
2 . Derschau, M. von (’ 09 ) : Beitragezur pflanzlichen Mitose : Centren, Blepkaroplasten. Pringsb. 
Jahrb. wiss. Bot., Leipzig, xlvi, 1. Heft, pp. 103-17. 
3 . Digby, L. (’ 09 ) : Observations on Chromatin Bodies. Ann. of Bot., xxiii, No. 91, July, pp. 
49 I ~ 5 ° 2 . 
4. Farmer, J. B., and Moore, J. E. S. (’ 05 ) : On the Meiotic Phase (Reduction Divisions) in 
Animals and Plants. Quart. Journ. Micr. Sc., xlviii, Part 4, Feb., pp. 489-557. 
5. Farmer, J. B., and Shove, D. (’ 05 ) : On the Structure and Development of the Somatic and 
Heterotype Chromosomes of Tradescantia virginica. Quart. Journ. Micr. Sc., xlviii, 
Part 4, Feb., pp. 559-69. 
6. Flemming, W. (’ 87 ) : Neue Beitrage zur Kenntniss der Zelle. Archiv fur mikroskop. Anat., 
Bonn, xxix, pp. 389-463. 
7. Goldschmidt, R. (’ 05 ): Eireifung, Befruchtung und Embryonalentwicklung des Zoogonus mirus , 
Lss. Zool. Jahrb., Jena, xxi, pp. 607-54. 
8. -(’ 08 ) : Die Chromatinreifung der Geschlechtszellen des Zoogonus mirus , Lss., 
und der Primartypus der Reduktion. Arch. f. Zellforschung, Leipzig, ii, 2. Heft, Dec., 
pp. 348-70* 
9 . Gregoire, V. (’ 06 ): L’Eldment chromosomique dans les cellules vegdtales. La Cellule, 
xxiii, 2 e fasc., pp. 311-53. 
10. -(’ 07 ): La Formation des Gemini hdt^rotypiques. La Cellule, xxiv, 2 e fasc., 
pp. 369-420. 
11 . - (’ 10 ): Les Cineses de maturation dans les Deux Regnes (2 e Memoire). La 
Cellule, xxvi, 2 e fasc.,pp. 223-422. 
12 . Hacker, V. (’ 07 ) : Die Chromosomen als angenommene Vererbungstrager. Ergebn. Fortschr. 
Zook, Jena, i, 1. Heft, pp. 1-136. 
13 . Hyde, Edith (’ 09 ): The Reduction Division in the Anthers of Hyacintkus orientalis. Con¬ 
tributions from the Bot. Lab. of Ohio State University, xlvii, June, pp. 539 ~ 44 * 
