338 
Journal of Agricultural Research 
Vol. XXIII, No. s 
of water content simultaneous with a deposit of starch, the latter 
beginning the fourth day after flowering. 
During the second stage there is little change in water content, but 
there is a heavy increase in starch content—that is, there is a deposit 
of starch which is not displacing water. The volume of the kernel 
/ r LOH'£’/?/A/e /&20 
5 * Average total water content of kernels of Jet barley from flowering to maturity at Aberdeen, 
Idaho, in 1920. 
continues to increase during this period. The increase in volume must 
come either from the increase in number of cells in the endosperm or 
from the increase in size of cells already formed. Most of the expansion 
is probably of the latter nature. The increase in volume is quite evident 
in figures i, 2, and 3, which show the length, lateral diameter, and dorso- 
ventral diameter by days for both varieties. 
The third stage, beginning about the twenty-second day in the Jet 
variety, shows a decided loss of water. While the rate of starch deposit 
