110 
Psyche 
[Vol. 87 
son with meganeurids suggests that the complete wing of analis was 
probably about 200 mm long, indicating a wing expanse of about 
420 mm. 
I have identified the type specimen as a fore wing on the basis of 
the slope of the hind margin, the hind wing of protodonates having 
a broad anal area basally. However, the basal part of the wing of 
analis! widens more abruptly than that of the fore wing of other 
protodonates so far known, although the base itself is narrow. 
As mentioned above, I have placed analis in the genus Palaeo- 
therates in preference to naming a new genus and for the present I 
consider the shape, size and nature of the anal vein to be specific 
characteristics. The species is noteworthy as the sixth specimen of 
Protodonata to be found in the Upper Carboniferous of North 
America, and as the largest species of that series of fossils. 
Order Caloneurodea 
This taxon, chiefly based on the genus Caloneura from the Upper 
Carboniferous of Commentry, was originally named by Handlirsch 
(1929) as a suborder of the order Protorthoptera. Martynov (1938), 
after studying a series of related genera from the Permian of the 
Soviet Union, elevated the group to an order. Since then, the 
Commentry species have been revised (Carpenter, 1961) and many 
other genera and families have been named to receive species from 
the Permian of the Soviet Union and the Permian and Upper 
Carboniferous of the United States (Tillyard, 1937; Carpenter, 1943, 
1970). Some of the additional species have led to a broadening of 
the concepts of the order and its relationships. These general topics 
will be considered in another article dealing with Caloneurodea 
from the Permian of Oklahoma, but I include here a brief statement 
of the characteristics of the order, as a preface to the description of a 
new species contained in the Klose collection and apparently 
belonging to the order. 
Although some of the body structures are known, the order 
Caloneurodea is based mainly on wings, which show three major 
characteristics: (1) the homonomous nature of the fore and hind 
wings, the hind pair lacking an enlarged anal area; (2) the origin of 
CUA from M near the wing base and its proximity to CUP for most 
of its length; (3) the numerous cross veins, which are either straight 
or somewhat reticulate and are very strongly developed and thick, 
