1980] 
Cady, Tietjen, & Uetz — Schizocosa 
233 
woods were found in leaf litter (depth 4-6 cm) and under a 
herbaceous “microcanopy” immediately adjacent to a spring seepage, 
while the spiders along Aspey’s ecotone were in a depressed area 
where leaf litter and moisture collected. The greatest number of 
spiders in the present study were found in areas with similar 
characteristics. Such microhabitats provide both soil moisture and 
protective cover (i.e., herbaceous growth and leaf litter). In addition, 
deep litter may also provide the high prey density needed for egg 
production by females (Uetz 1975, 1979). 
Tietjen (1979) suggested that airborne pheromones may be involved 
in attracting S’, ocreata to particular areas for courtship and 
copulation. Such pheromones might be expected to collect under the 
leaves occupied by the relatively sendentary females. This may 
explain the denser concentrations of this spider and the social 
attractions Aspey (1976) proposed. We therefore agree with Aspey’s 
premises that microclimate and possible social attractions are 
determining factors for S. ocreata’s pattern of local distribution; 
however these microclimates or areas of congregation are not unique 
to ecotones, and S. ocreata should not be termed an “edge” species. 
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