1979] Evans, Kug/er & Brown 
Scolebythus madecassus 
47 
09551 and Oeser (1961). but other commonly used synonyms are 
given as well. Numbers of abdominal, rather than metasoma' 
segments are used throughout. ’ 
Apparatus not completely internalized; sides of sternum 7 large 
enclosmg all but a small triangle of tergum 8 and tips of thfrd 
valvulae and aculeus (Fig. 1). Eighth tergum (spiracular plate) (Fig 
(subcorneal in dorsal view, not reduced mesad, with heavy 
antecostal ridge that terminates anterolaterally in long acute nro 
cesses Minth tergum (quadrate plates) (Fig. 3) completely divided 
sd'erot z Y ed y m ^ brane ’ narr ° W (max W/max L = 35%) moderately 
sclerotized, apodeme prominent only in upper half of length No 
ox e e 8HF ^ Plate i FirSt V3lviferS ‘‘“"gutar P> a tes gono- 
ram s h 3 att3Ched t0 near the a P ex of:first 
ramus but slung beneath it on a narrow stalk. First valvulae 
(lancets, gonapophyses 8) (Fig. 3) long, very slender, acute; each 
with a large proximal valve and a smaller, weaker distal valve- apex 
with a single small barb and 3 tiny denticles. Dorsal lamina (above 
groove) sclerotized only in distal third of length from valves. Second 
alvifers (oblong plates, gonocoxites 9) (Fig. 3) hinged between the 
dorsal and ventral arms. Dorsal arm long and narrow, its ventral 
wlfh f 3 rad ' n / ' nt t ° m h en \ brane . ce Phalad; intervalv.fer articulations 
with 23 and 27 tncho.d sensilla. Ventral arm curved and evenly 
trichoid t0 artlCUlar P rocess of sting; second rami with 63 and 67 
trichoid sensilla present on all but extreme ends. Second valvulae 
(sting gonapophyses 9) (Fig. 3) moderately arched so that tip is 
directed obliquely ventrad. Profile tapers evenly to tip which is 
about twice as wide as high and is free of ridges and barbs. Placoid 
or coelocomc sensilla present along caudal 45% of length, increas- 
mgly dense ap,cad Sting base truncate, with rounded shoulders 
when seen from below (F,g. 5), not arched when viewed from the 
ont. Articular processes long, slender and curved. Furcula (Figs 3 
T ral a ™ S that d ° n0t curve over sides of sting 
ba e. Third valvulae (gonostyli) (Figs. 3, 6) 2-segmented, both parts 
ell sclerotized and with numerous trichoid and basiconic sensilla 
The morphology of the Scolebythus madecassus sting apparatus 
supports the close association of the Scolebythidae with the Bethyli- 
dae (Evans, 1963; Brothers, 1975), rather than with Sapygidae or 
Anthoboscnae (Evans, 1963). The presence of a furcula and 
post.nc.sion (incisura postarticularis of Oeser), and lack of socii 
(pygostyles, cere) separates Scolebythus from the terebrantes 
ough sincids and some proctotrupids lack socii) (Oeser, 1961 
