74 
Psyche 
[March 
year for army ant nuptial flight activity in the region of Barro 
Colorado Island’—is in accord with my data, and indicative of this 
interface being in fact a wet sunny period. 
The fauna of Legon is a complex mix of forest relicts, secondary 
successional species and true savanna forms: investigations have 
covered such diverse groups as snakes, ponerine ants, hemipters and 
mantises (Leston, 1972, 1979). In practice it is seldom difficult to 
categorize a particular animal by habitat. The relative numbers, in a 
degraded forest habitat, is a function of the degree of deforesta¬ 
tion-most species are not directly climate-limited—modified by the 
varying abilities of savanna taxa to saltate (Leston, 1979). 
The traps reported on by Haddow et al. (1966) covered both 
tropical forest and degraded forest but unfortunately the results are 
given in pooled form, making a numerical habitat comparison diffi¬ 
cult: the pooling of data from three traps run at different times of 
the year makes an investigation of periodicities equally impossible. 
But overall, despite species differences—which probably reflect an 
oversplitting taxonomy—there is a marked similarity in the dory¬ 
lines trapped at Legon, on the edge of the Guinea forest bloc, and 
that of Entebbe, 3,600 km distant on the edge of the Congo forest 
bloc. 
As at Legon, the most abundant species at Entebbe was an Alao- 
pone: these are hypogeic ants of degraded forest habitats. Typhlo- 
pone fulvus, assumed by Haddow et al. (1966) and Wheeler (1922) 
to be hypogeic, was also numerous at both localities: my field obser¬ 
vations show it frequently forages at the surface in forest and 
degraded forest areas. Some Dorylus sensu stricto were frequent at 
both sampled places: probably for the most part savanna ants which 
invade degraded forest. Anomma species were relatively infrequent 
at both too: as noted above they are essentially genuine forest 
dwellers, surface foraging, the ‘notorious driver ant’ (to cite Had¬ 
dow et al.). Rhogmus fimbriatus, about as frequent at Entebbe as at 
Legon, is a hypogeic, mainly savanna, species. 
The similarity of the two faunas over an extensive transcontinen¬ 
tal area suggests a widespread pattern of resource partitioning but 
why the alates should space themselves out in male flights through 
the hours from dusk to dawn, as Haddow and his colleagues 
indicate—found too in the American Ecitonini (Kannowski, 1969)— 
is less easy to interpret in such terms. And the most remarkable 
feature of the Legon results, the synchrony of the species’ cycles, 
