1979] 
Carpenter — Permian Insects from Oklahoma 
285 
Figure 16. Moravia grandis, n.sp. Photograph of holotype, MCZ 8647a; maxi¬ 
mum length of preserved part of wing, 74 mm. Permian of Oklahoma. 
gens the intercalary veins were also confined to the branches of RS 
and MP but were very irregular and formed by alignment of the 
sides of the two rows of cells bordering the main veins. In Carrizop- 
tera arroyo they are so short and irregular as to be hardly recogniza¬ 
ble. In the new species, Moravia grandis, described below, the radial 
sector is not preserved, but the intercalary veins are not present 
between the branches of MA, MP, or CUA, the spaces between the 
veins being filled with a reticulate archedictyon. The nature of the 
intercalary veins in the Calvertiellidae, therefore, would seem to 
suggest that these veins developed in that family quite independently 
of their occurrence in other orders or families. They appear to have 
arisen by the alignment of the sides of the cells forming the archedic¬ 
tyon between the main veins. The development of such intercalary 
veins, in association with the increased fluting along the veins, may 
have provided more support for the wing membrane than the origi¬ 
nal archedictyon. In any case, I believe that the presence or absence 
of the intercalary veins should not be given very much weight in 
considering evolutionary relationships. 
