318 
Psyche 
[December 
dibular prey carriage over the ground also occurs in both of these 
families, and the manner of dragging the prey backward (Fig. 2a) is 
remarkably like that of many Pompilidae. Williams was, however, 
correct in characterizing carriage such as that shown in Figure 2b as 
unique. 
Members of other genera of Bethylidae are known to carry their 
prey to a second site for oviposition after being stung; this includes 
Bethylus, Holepyris, Parascleroderma, and to a certain extent Alle- 
pyris (Yamada, 1955). However, only Epyris actually constructs a 
nest. Further work regarding the comparative ethology of Epyris, as 
well as other members of this family should prove very rewarding. 
Acknowledgments 
This work was partially supported by National Science Founda¬ 
tion grants DEB75-17142 and BNS78-17818 to Howard E. Evans. 
We thank T. J. Spilman of the USDA Systemic Entomology Labora¬ 
tory for identifying the tenebrionid larvae. 
Literature Cited 
Bridwell, J. C. 
1917. A note on Epyris (Hymenoptera, Bethylidae) and its prey. Proc. Haw. 
Ent. Soc. III(4):262 263. 
1919. Some notes on Hawaiian and other Bethylidae (Hymenoptera) with the 
description of a new genus and species. 2nd paper. Proc. Haw. Ent. Soc. 
IV(2):291-314. 
Evans, H. E. 
1964. A Synopsis of the American Bethylidae (Hymenoptera, Aculeata). Bull. 
Mus. Comp. Zool. 132(1): 1-222. 
1974. Digger wasps as colonizers of new habitat. J. N.Y. Ent. Soc. 82(4):259—67. 
1978. The Bethylidae of America north of Mexico. Mem. Am. Ent. Inst. No. 27. 
332 pp. 
Gordh, G. 
1976. Goniozus gallicola Fouts, a parasite of moth larvae, with notes on other 
bethylids: (Hymenoptera: Bethylidae, Lepidoptera, Gelechiidae). 
U.S.D.A. Tech. Bull. 1524, 27 pp. 
Gordh, G. and H. E. Evans 
1976. A new species of Goniozus imported into California from Ethiopia for the 
biological control of pink bollworm and some notes on the taxonomic 
status of Parasierola and Goniozus (Hymenoptera: Bethylidae). Proc. 
Ent. Soc. Wash. 78(4):479-89. 
Malyshev, S. 1. 
1968. Genesis of the Hymenoptera and the Phases of their Evolution. Methuen 
& Co., Ltd., London. 319 pp. 
