1979] 
North & Shaw — Pterophylla camellifolia 
367 
Morphological characters were determined from 13 previously 
collected specimens and from 192 males collected from 15 different 
colonies in 1977 and 1978. We examined characters used previously: 
length of body, length of metathoracic femur, length and width of 
tegmina, length and width of subgenital plate, length of the subgeni¬ 
tal plate slit, and shape of cerci (Beutenmuller 1894, Caudell 1906, 
Blatchley 1903, 1920, Walden 1911,Hebard 1941, Shaw and Carlson 
1969). In addition we measured the distance from the frons to the 
distal end of the tegmina. Measurements were made with calipers and 
an ocular micrometer. 
Results 
In Iowa, P. camellifolia is restricted to gallery forests along major 
rivers, and colonies in central Iowa are disjunct from those of eastern 
Iowa (150 to 200 km). In western Iowa, distribution of P. camellifolia 
is limited to two colonies, one in Waubonsie State Park in Fremont 
County (colony number 10), and another in a county park in Page 
County (colony number 11) (Fig. 1). 
Table 1. Nested ANOVA. A = Subgenital plate length. B = Subgenital plate slit 
length. C = Pulse number per chirp. 
% of total 
Source 
d.f. 
MS 
variation 
A. Colonies 
14 
34.33 
87.33 
Individuals (Colonies) 
190 
0.37 
12.67 
B. Colonies 
14 
9.36 
76.99 
Individuals 
190 
0.21 
23.01 
C. Colonies 
19 
1976.53 
79.50 
Individuals (Colonies) 
191 
34.39 
14.52 
Records (Individuals Colonies) 
4009 
0.69 
5.98 
All characters showed significant F values (ranging from 2.54 to 
74.02, p < .01). However, of the 11 morphological and behavioral 
parameters measured, only subgenital plate length (SPL), subgenital 
plate slit length (SPSL) and number of pulses per chirp (P/ C) showed 
greater variation between colonies than within colonies (Table 1). 
Also these three characters were the only ones by which the colonies 
were separated into two significantly different populations by the 
Duncan’s multiple range test (Table 2). SPSL is not shown in Table 2 
