1979] 
North & Shaw — Pterophylla camellifolia 
371 
Fig. 2. Ventral arm of male cercus, ventral view. A. Northern population, colony 
#10. B. Western population, colony #3. C. Western population, colony#4. D. Western 
population, colony #5. 
extreme cold. Such a brood-splitting could have resulted in brood III 
(central and southeast Iowa) and brood IV (southwest Iowa and 
south to Texas). 
Smith (1957) takes a position intermediate to that of Deevy (1949) 
and that proposed by Transeau (1941), Braun (1951), Thomas (1951), 
and Alexander and Moore (1962). Smith’s evidence is principally 
southern disjunct, relict populations of boreal vertebrates whose 
main contemporary distributions lie “partially or wholly within gla¬ 
ciated North America”. These relict populations, widely separated 
from the main segment of their species ranges, occur in such states as 
Missouri, Arkansas, Kansas, Tennessee, and Georgia. 
Such western (Missouri, Arkansas, Kansas) and eastern (Tennes¬ 
see, Georgia) separation of a preglacial population of P. camellifolia 
could have been responsible for differentiation of the western and 
