D.:?iLGFor ih riVi DULTS ahd c*:>als 
i *, 'i :.n 
C - 
?, i : d :. ,:• tr ;- , L . 
A LIS dKTATZOS 
I ub ittod to the Faculty 
of the 0£d<m Graduate chool o* Science 
in Candid&cy for the 4>ogvoo of 
Doctor o ' Philosophy# 
fJKPUH £<T Of- ZOOLOGY 
rhor:uj buue -iaaonpotte 
J .4 J ■ si 
Xf£3 
