284 MOULTONIA : A NEW BORNEAN GESNERACEOUS GENUS. 
As an intermediate condition we may bring into the 
case the evidence offered by Monophyllcea. 
We have pointed out that Moultonia is allied to Mono - 
phyllcea. This genus we only know from the description 
and figure given by Clarke.* Clarke suggests, we think 
rightly, that the folium unicum of his plant is a cotyledon. 
There appears to be in Monophyllcea a single stalked leaf— 
to speak conventionally—as in Moultonia , but the stalk is 
shorter and at the junction of lamina and stalk scapose 
inflorescences arise. At the summit of the scapes the 
flowers are disposed in a unilateral, spicate raceme which 
curls over circinately. We should interpret all this con¬ 
struction as that of a hypocotyl elongated in the form of a 
petiole and bearing at its extremity a cotyledonary lamina. 
We should expect in the young protocorm to find a second 
smaller arrested cotyledon. At the point where cotyledonary 
lamina and hypocotyl join, the inflorescence arises from the 
hypocotyl. The meristem for the inflorescence is localised 
at this region. 
Now Monophyllcea is of special interest to residents in 
Borneo, as that country contains at least three of its 
known species—some ten in number. The study of its 
morphology from the seedling stage would be full of value 
and would throw much light on the problems raised by 
Moultonia. We therefore hope that those interested will 
try to secure specimens and seeds of both genera. Mono - 
phyllcea can be recognized among the Gesnerads by its 
single large leaf, presenting a similar appearance to that of 
Moultonia but with the inflorescences, as indicated above, 
distinct from the leaf-stalk and midrib. 
Of the allied Epithema we have insufficient knowledge 
to allow of our making a suggestion of interpretation. 
The following morphological series within the plants 
named issues from what we have said:— 
Streptocarpus polyanthus , Hook.—Hypocotyl not elon¬ 
gated, cotyledon sessile, scapose inflorescence with 
open biparous branching developing from hypocotyl 
base of cotyledonar lamina. 
Streptocarpus Rexii, LindL—Hypocotyl shortly elon¬ 
gated, giving apparent stalk to cotyledon, scapose 
* C. B. Clarke, Cyrtandreo, in De Candolle Monogr. Phanerog. v. (1883- 
87), 181, t. xx. 
