BY PROF. BAYLEY BALFOUR AND W. W. SMITH. 
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inflorescence (one flower) developing from hypocotyl 
at base of cotyledonar lamina. 
Monophyllcea. —Hypocotyl much elongated, giving ap¬ 
parent long stalk to cotyledon, scapose inflorescence 
with unilateral racemes developing from hypocotyl at 
base of cotyledonar lamina. 
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Moultonia. —Hypocotyl very much elongated, giving 
apparent very long stalk to cotyledon, inflorescence 
disposed in umbels originating along middle line on 
upper surface of whole length of elongated hypocotyl 
and on midrib of lamina (but not showing special 
relation to the primary veins). 
We do not overlook other explanations that might be 
given of the construction in Moultonia . Thus, in absence 
of all evidence of the seedling condition, it might be held 
that in both Moultonia and Monophyllcea the folium unicum 
is really an epicotylar leaf with which the inflorescence is 
more or less “ congenitally concrescent.” In such a view, 
the unilateral disposition of the flowers on the scapes in 
Monophyllcea might be regarded as a stage towards the 
complete fusion of scape with leaf in Moultonia. To us 
such concrescences do not appeal. 
A more illuminative comparison may be made with what 
is seen in Chirita hamosa, K. Br., of which, however, we 
have not yet the clear explanation. In Plate III. is a 
figure of this plant when in flower. The opposite leaves 
are petiolate, and the flowers arise in a line upon the upper 
surface of each petiole. They are epipetiolar. The buds 
on the petiole are not, however, all flower-buds ; some are 
foliage. The sequence is irregular. The position of the 
inflorescence partially recalls that of Moultonia. There is 
no concrescence here; simply foliar evolution of flower 
and buds. The seedling of Chirita hamosa has cotyledons 
differing in size and separated by a hypocotylar elongation. 
The upper is the larger, becomes stalked, and has quite 
the form of the adult leaf, but it never bears flowers or 
buds. Possibly, then, what we have been describing in 
Moultonia as a protocorm outgrowth may be after all an 
epicotylar leaf with epiphyllous inflorescences more ex¬ 
tended than in Chirita hamosa. 
The flower-structure of Moultonia is not without special 
interest. The gynaeceum is closed at the top by a solid 
cone like a style supported upon a stylopod, and this seems 
