Hill, Johnson and Blaxell, Systematic studies in the eucalypts 
261 
regularly reticulate, irregularly circumferentially winged or flanged, more or less 
pyramidal or flattened-pyramidal, with winged ribs radiating from the paler, ventral 
hilum; stomata large and translucent, giving a bluish sheen to the leaves. 
Two subseries are distinguishable within the series (Table 2). Subseries Incmssatosae is 
made up of a number of species that have all been included in a broad application of 
E. incrassata at some time or other. Several of these have already been clarified (Table 
3, and see Chippendale 1988, Brooker & Hopper 1993), and others are discussed below. 
The subseries is widely distributed across the 'mallee country' of southern Australia, 
from south-western Western Australia east to south-western New South Wales and 
western Victoria (Figs. 2,3) This account and classification summarises our revisionary 
studies of this series, although further study of the variation in, and geographic extent 
of, E. captiosa Brooker & Hopper is still required. A tabular comparison of characters is 
provided to facilitate identification within the series, rather than a dichotomous key 
(Table 4). 
Table 2. A dassification of Series Incrassatae 
Series Incrassatae 
Subseries Incrassatosae (£. incrassata sens, lat.) 
E. captiosa 
E. incrassata 
E. angulosa 
E. ceratocorys 
E. costa ta 
subsp. murrayana 
subsp. costata 
E. capitanea 
E. grossifolia 
E. singularis 
Subseries Tetrapterosae 
E. tetraptera 
E. stoatei 
E. forrestiana 
E. dolichorhyncha 
Table 3. Published names in Series Incrassatae (accepted names in Bold type, taxonomic 
synomyms represented by nomenclatural synonyms represented by ='). 
E. incrassata Labill., Nov. Holl. PI. Sp. 2: 12 (1806). 
E. angulosa Schauer in Walp., Repert. Bot. Syst. 2: 925 (1843). 
E. cuspidata Turcz., Bull. Soc. Nat Mosc. 22(2): 22 (1849); = E. angulosa. 
E. tetraptera Turcz., Bull. Soc. Nat. Mosc. 22(2): 22 (1849). 
E. acutangula Turcz., Bull. Cl. Phys.-Math. Acad. Imp. Sci. Saint-Petersbourg 10: 337 (1852); = 
E. tetraptera. 
E. costata F. Muell., Trans. Victorian Inst., 33 (1855). 
£. forrestiana Diels in Diels & Pritzel, Bot. Jahrb. Syst. 35: 439 (1904). 
E. angulosa Schauer var. ceratocorys Blakely, Key Eucalypts, 174 (1934); = E. ceratocorys. 
E. stoatei C. Gardner, J. & Proc. Roy. Soc. Western Australia 22: 126 (1936). 
E. forrestiana subsp. dolichorhyncha Brooker, J. & Proc. Roy. Soc. Western Australia 56(3): 74 (1974); = 
E. dolichorhyncha. 
E. ceratocorys (Blakely) L.A.S. Johnson & K.D. Hill, FI. Australia 19: 508 (1988). 
E. dolichorhyncha (Brooker) Brooker & Hopper, Nuytsia 9: 57 (1993). 
E. captiosa Brooker & Hopper, Nuytsia 9: 55 (1993). 
