266 
Telopea 9(2): 2001 
E. incrassata as a local variant of a highly variable species. Following more detailed 
studies, we now consider that a number of taxa can be distinguished within 
E. incrassata sens, lat., of which £. angulosa is one (Table 2). Chippendale (1988) applies 
essentially the same circumscription of E. angulosa in his treatment. 
Mallee to 6 m tall. Bark smooth, grey to grey-brown, shedding in ribbons. Juvenile 
leaves disjunct, broad-elliptical, to 10 cm long, 7 cm wide. Adult leaves disjunct; 
broad-elliptical, apiculate, glossy bright green, 8.0-14.0 cm long, 25-50 mm wide; 
petioles terete, 15-30 mm long; lateral veins at 35^15° to midrib, moderately spaced, ± 
degenerate; oil glands dense and distinct; intramarginal vein 2-4 mm from margin. 
Umbellasters axillary, 3-7-flowered; peduncles flattened, broadly apically expanded, 
10-20 nun long; pedicels terete to quadrangular, 3-5 mm long. Mature buds ovoid, 
coarsely distinctly costate, 11-17 mm long, 7-9 mm diam.; calyptra conical, usually not 
beaked, c. Vi as long to as long as hypanthium. Fruits cup-shaped, distinctly broadly 
costate, 3-4-locular, 15-25 mm long, 15-25 mm diam.; calyptra scar and stemonophore 
flat, c. 1-2 mm wide; disc steeply depressed, 2 -5 mm wide; valves enclosed. Seeds ± 
pyramidal, deeply and regularly pitted, not winged, black, 2-3 mm long; hilum 
ventral; chaff red brown, smaller, angular (Fig. lc, d). 
£. angulosa is a coastal taxon, occurring in dense shrub thickets, usually on calcareous 
substrates, along the southern coast of Western Australia from West Cape Howe east 
past Israelite Bay, and in southern Eyre and Yorke Peninsulas and Kangaroo Island in 
South Australia (Fig. 2). It is distinguished within the series by the coarse leaves, buds 
and fruits (see also Table 4). 
Selected specimens (from 69 examined): South Australia: Encounter Bay, Cleland 16, Feb 1921 
(NSW); between Fishery Bay and Cape Wiles, Blaxell 2063 & Johnson, 17 June 1983 (NSW, CANB); 
Stamford Hill, Maiden s.n., Jan 1907 (NSW); Port Lincoln to Coffins Bay, Maiden s.n., Jan 1907 (NSW); 
Mt Hope, Noble 7 & Bazvden, Feb 1981 (NSW); 4 km S Mt Hope Post Office 85 km NW Port Lincoln, 
Pickard 2522, 10 Dec 1973 (NSW); Porter Bay, S of Port Lincoln, Erosion control area between Bay 
and Point, Symon 13712, 20 Dec 1983 (AD, CANB, NSW, US). 
117' 
122 ' 
127' 
(W.A. occurrences) 
© E. captiosa 
© E. ceratocorys 
28‘ 
117* 
122 ' 
® E. grossifolia 
• E. incrassata 
A E. singularis 
Fig. 2. Distribution of E. angulosa p.p., E. captiosa, E. ceratocorys, E. grossifolia, E. incrassata, E. singularis. 
