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2:1); fruits shallowly ribbed to almost smooth, urceolate, medium size (9-16 mm long, 7— 
21 mm diam., see also Table 4). 
Two subspecies are recognised, as follows 
3A. Eucalyptus costata F. Muell. subsp. costata 
Fruits ovoid to cup-shaped, large (11-16 mm long, 9-12 mm diam.), distinctly ribbed, 
valves deeply enclosed to slightly below rim level (Fig. li, j). 
Restricted to South Australia, on red sand or sandy loam, central Eyre Peninsula, 
Yorke Peninsula and the upper and lower Murray Mallee regions, east almost to the 
Victorian border (Fig. 3). 
Intergradation occurs in marginal regions with E. angulosa and hybrids are known 
with E. rugosa R. Br. ex Blakely. 
Conservation status: not considered to be at risk. 
Selected specimens (from 57 examined): South Australia: c. 10 km N of Cummins, Blaxell 2062 & 
Johnson, 17 June 1983 (NSW, CANB); c. 16 km south of Curramulka, Blaylock 1801 ,30 Apr 1973 (AD, 
NSW); Goolwa, c. 65 km south-south-east of Adelaide, River Murray Lakes, Boomsma, 9 Nov 1967 
(AD, NSW); Padthaway, ca. 40 km SSW of Bordertown, Canty, 23 Oct 1980 (AD, NSW); Mt 
Weedind, about 15 miles [24 km] South from the Gawler Ranges and 125 miles [200 km] a little 
west of north from Port Lincoln, Gill, Dec 1912 (NSW); 74 km from Whyalla on Cowell road, Hill 
2145 & Johnson, 1 Nov 1986 (NSW, AD, CANB, MEL, PERTH); 7 km north of Arno Bay on Lincoln 
Highway, Hopton 21 ,21 Oct 1988 (AD, NSW); c. 17 km S of Bordertown on Naracoorte road, Johnson 
7920 & Wilson, 20 Feb 1975 (NSW); 7 miles [11 km] south of Monarto South, McKern, 24 Aug 1958 
(NSW); Murray Bridge, Maiden, Jan 1907 (NSW); 9 miles [14 km] from Port Broughton, towards 
Bute, Phillips, 11 Nov 1962 (CANB, NSW); 23 miles [36.8 km] from Bordertown, towards Keith, 
Phillips, 21 Oct 1966 (CANB, NSW); stock route 14 km SW Buckleboo HS [homestead], 150 km 
WNW Port Augusta, Pickard 2496 ,9 Dec 1973 (NSW); between Bordertown and Keith, 8 miles (13 km) 
WNW of Bordertown, Smith 67/201 ,17 Nov 1967 (MEL, NSW); area between Oak Dam (Frenchs 
Dam) and Gypsum Hills c. 32 miles [51 km] N of Overland Comer, Symon 3631, 9 Oct 1965 (AD, 
Fig. 3. Distribution of E. angulosa, E. capitanea, E. costata subsp. costata, E. costata subsp. murrayana. 
