Hill, Johnson and Blaxell, Systematic studies in the eucalypts 
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hypanthium. Fruits urceolate, distinctly costate, 3-4-locular, 15-22 mm long, 12-15 mm 
diam.; calyptra scar and stemonophore raised, 1-2 mm wide; disc steeply depressed, 
3-4 mm wide; valves deeply enclosed. Seeds ± pyramidal, deeply and regularly pitted, 
narrowly winged circumferentially and with ventral ridges, dark grey-brown to black, 
3-4 mm long; hilum ventral; chaff red brown, smaller, angular. (Fig. lg, h). 
E. capitanea is distinguished within the E. incmssata complex by the following 
combination of characters: mallee; calyptra short (l:b < 2:1); fruits ribbed, urceolate, 
large (15-22 mm long, 12-15 mm diam.). See also Table 4. 
Locally frequent but at present known from one site only, on the crests of red desert 
dunes near Koonibba (Fig. 3). Associated taxa include E. yumbarrana Boomsma and 
Triodia, with E. oleosa subsp. repleta L.A.S. Johnson & K.D. Hill and £. brachycalyx 
Blakely lower on dune slopes and on swales. 
Conservation status: a species of restricted occurrence, although the extent is not 
known (2K). 
The epithet is from the Latin capitaneus, head or chief, in reference to the larger size of 
this species in all parts in comparison to related taxa. 
Specimens examined: South Australia: 8.4 km N of Koonibba towards Yumbarra Conservation 
Park, Brooker 8466 ,9 Mar 1984 (CANB, NSW); c. 2 km NW of Koonibba Hill on access track to new 
wheatlands, c. 30 km E of Penong, Hill 714 & Blaxell, 15 Nov 1983 (NSW, AD, CANB, PERTH); 7.3 km 
N of highway toN of Charra turnoff (30 km E of Penong, 40 km W of Ceduna), Hill 2157 & Johnson, 
2 Nov 1986 (NSW, AD, CANB, MEL, PERTH). 
5. Eucalyptus grossifolia L.A.S. Johnson & K.D. Hill, sp. nov. 
E. angulosam plusminusve approximans sed habitu erecto, sparso, emergenti nec 
humile nec denso differt. 
Type: Western Australia: 1.3 km from Kambellup on Woogenellup rd (34°'S 117°59'E), 
K.D. Hill 2444, L.A.S. Johnson & D.F. Blaxell, 12 Nov 1986 (holo NSW; iso PERTH). 
Mallee to 6 m tall. Bark smooth, grey to grey-brown, shedding in ribbons. Juvenile 
leaves disjunct, ovate to elliptical, to 10 cm long, 5 cm wide. Adult leaves disjunct; 
lanceolate to broad-lanceolate or narrow-elliptical to ovate, acuminate, glossy bright 
green, 5.0-12.0 cm long, 20-50 mm wide; petioles flattened, 11-26 mm long; lateral 
veins at 35-45° to midrib, moderately spaced, ± degenerate; oil glands dense and 
distinct; intramarginal vein 1-2 mm from margin. Umbellasters axillary, 7-flowered; 
peduncles flattened, apically expanded, 12—18 mm long, to 10 mm wide apically; 
pedicels terete, 0-4 mm long. Mature buds ovoid, sessile or shortly pedicellate, 
distinctly irregularly costate, 9-17 mm long, 6-8 mm diam.; calyptra conical, often 
beaked, V 3 -V 2 as long to as long as hypanthium. Fruits cup-shaped to obconical, 
distinctly irregularly costate, 3-4-locular, 10-20 mm long, 9-15 mm diam.; calyptra scar 
and stemonophore flat, c. 1 mm wide; disc steeply depressed, 2-4 mm wide; valves 
enclosed. Seeds ± pyramidal to patelliform, deeply and regularly pitted, narrowly 
winged circumferentially and with ventral ridges, dark grey-brown to black, 2-3 mm 
long; hilum ventral; chaff brown, smaller, angular (Fig. 4). 
E. grossifolia is most similar to E. angulosa, differing mainly in the slender, sparse, erect, 
emergent habit in contrast to the low, dense, bushy habit of the latter (see also Table 4). 
E. angulosa is also largely restricted to calcareous sand deposits on or near the coast, 
whereas E. grossifolia occurs on lateritic residuals, often with a shallow siliceous sand 
mantle, some distance from the coast. 
A component of rich mixed mallee shrublands, associated species including E. uncinata 
Turcz., E.falcata Turcz., E. pleurocarpa Schauer, E. pachyloma Benth., E. tetraptera Turcz. 
