Archer, Phaeographis and Phaeographina 
467 
Thallus off-white to pale olive green, thin, corticolous, surface smooth and shiny; 
apothecia lirelliform, black, conspicuous, immersed or sometimes subsessiJe, straight, 
curved or sinuous, sometimes branched, 1-4 mm long, 0.2-0.5 mm wide, often in 
substellate clusters to 4 mm wide, with a thin, white thalline margin; proper exciple 
thin, completely carbonised; hymenium (75-)100-125 pm tall; disc matt black, 
epruinose; ascospores 8 per ascus, irregularly 2-seriate, very pale brown, 
(15-)17-21(-23) pm long, 6-7 pm wide, 4-locular. 
Chemistry: no compounds found. 
Also examined: Queensland: New England Highway, 20 km SW of Yarraman, Haffellner 19362, Sep 
1986 (GZU); Conors Range, c. 20 km SW of Sarina, Lumbsch 10983d, p.p., Oct 1994 (ESS); Bowenia 
State Forest, Stony Creek, 25 km NNW of Yeppoon, Elix 34571, Aug 1993 (CANB); Forrest Beach, 
16 km SE of Ingham, Streinmnn 28849, June 1984 (CANB); Mcllwraith Range, 29 km NE of Coen, 
Streitnann 56831, Oct 1995 (CANB). 
New South Wales: Ku-ring-gai Chase National Park, the Basin camping area, on felled tree. Archer 
G261, Sep 1998, (NSW). 
Phaeographis elaeina is characterised by the immersed, relatively wide, black lirellae, the 
4-locular ascospores and the thin, completely carbonised proper exciple. This latter 
feature distinguishes the species from the somewhat similar P. subintricata, which has 
a thin carbonised proper exciple open at the base (laterally carbonised), and 
P. pseudomelana, which is apically carbonised. Phaeographis elaeina has been found in 
Queensland and New South Wales. Reported substrates include Casuarina and 
Excoecaria. 
Phaeographis eludens (Stirt.) Shirley (Fig. lc, 2c) 
(Shirley 1889: 197). 
Graphis eludens Stirt. 
(Stirton 1881: 72). 
Type: Queensland: sine loc., as 'saxicola', F. Bailey 287 (holo BRI). 
Thallus fawn to pale fawn, 1-2 mm thick, saxicolous, surface smooth and dull 
(superficially resembling fine velvet); apothecia lirelliform, black, numerous, 
inconspicuous, open, immersed, straight, curved or sinuous, often branched and in 
sub-stellate clusters; individual lirellae 0.5-2 mm long, 0.1-0.2 mm wide, lacking a 
thalline margin; proper exciple absent; hymenium (100—)125—175(—200) pm tall; 
ascospores 8/ascus, 2-3-seriate, pale brown, 12-15 pm long, 5-7 pm wide, 4-locular. 
Chemistry: no compounds found. 
Also examined: Queensland: sine loc., Shirley s.n., Jan 1888 (NSW); Burleigh Heads National Park, 
on basalt boulders, Elix 1093, Aug 1975 (CANB); Isla Gorge National Park, 27 km NNE of Taroom, 
on sandstone rocks, Elix 35177, Aug 1993 (CANB); Tabletop, Harvey Range, 39 km WSW of 
Townsville, Streimann 37185, 37192, Aug 1986 (CANB). 
Phaeographis eludens is characterised by the substrate preference, the thick thallus, the 
small 4-locular ascospores and the absence of lichen compounds. Stirton (1881) 
compared his new species with Graphis hypoglauca Krempelh. [Phaeographis hypoglauca 
(Krempelh.) Zahlbr.] but the ascospores in that species are larger, 18-28 x 8-11 pm 
(Singh & Awasthi 1979). 
The species is so far known only from Queensland where it occurs on basalt and 
sandstone. The species was also reported from Moggill and Mount Perry [Brisbane] 
(Shirley 1889). 
