470 
Telopea 8(4): 2000 
straight, curved or sinuous, often branched, 1-4 mm long, 0.2-0.5 mm wide, often with 
a conspicuous, white thalline margin, sometimes forming substellate clusters; proper 
exciple thin, laterally carbonised, open at the base; hymenium 100-125 pm tall; disc 
matt black, epruinose; ascospores 8 per ascus, irregular 2-seriate, very pale brown, 
(14-)17-21(-24) pm long, 6-8 pm wide, 4-locular. 
Chemistry: no compounds found. 
Also examined: Queensland: Upper Coomera, Wilson s.n., as 'Graphis innata' (NSW). 
New South Wales: Sydney, Mosman Bay, Wilson s.n., Sep 1897 (NSW); Urunga, Hungry Head, 
c. 25 km SSW of Coffs Harbour, Archer G 212, Apr 1998 (NSW). 
Victoria: Cunningham, Wilson s.n., no date (NSW L5082); Metung, Wilson 877a, Mar 1889 (NSW); 
Lake Tyers, Wilson, no date (NSW 180877); Metung, Wilson s.n., Mar 1889 (NSW); Gippsland, 7 km 
S of Bonang, Verdon 4169, Nov 1978 (CANB, H). 
New Zealand: sine loc., C. Knight s.n. [as Graphis In/poleuca, nom. nud.] (NSW 155588). 
Phaeographis subintricata is characterised by the black, open lirellae, 4-locular 
ascospores, the incompletely carbonised proper exciple and the absence of lichen 
compounds. It resembles P. dendritica but is distinguished from that species by the 
smaller, 4-locular ascospores and the absence of norstictic acid. Phaeographis 
subintricata closely resembles P. elaeina and is only distinguished from that species by 
the completely carbonised proper exciple in the latter species, and from 
P. pseudomelana by the apically carbonised proper exciple in that species. 
Graphis subintricata and Graphis subtricosa were published simultaneously (Knight 
1882) and as G. subintricata is the first Graphis species described (no. 5; G. subtricosa no. 
6) it is taken to be the earliest name. Phaeographis intuntescens and P. extenuata were also 
published together (Muller 1893, loc. cit.) and are based on the same specimen, Wilson 
877, but the two new species were differentiated on the basis of the thallus colour. 
Muller (loc. cit., 1895) noted that P. subintricata was close to P. extenuata. The four 
species listed lack any features to distinguish between them. 
Reported substrates include Acacia, Eucn/phia, Lomatia and Notelaea. 
Additional syntypes of P. subintricata are in G, M and WELT (Filson 1986) and as all of 
these have not been seen, no lectotype is selected here. 
Species of uncertain status 
Phaeographis mucronata (Stirt.) Zahlbr. 
(Zahlbruckner 1923: 382) 
Graphis mucronata Stirt. 
(Stirton 1876: 95). 
Lectotype (fide Rogers 1982): New South Wales: Riverina District, H. Paton s.n., 1875 
(BM, not seen). 
The type was not available for examination; see discussion above under Phaeographis 
australiensis. 
