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Telopea Vol. 6(4): 1996 
Danthonia penicillata (Labill.) Beauv. van pallida (R. Br.) F. Muell. ex Maiden and 
Betche, Census New South Wales Plants: 29 (1922). 
Type: R. Brozvn '6232' (BM, holo, K!). 
Joycea clelandii (Vickery) Linder, comb. nov. 
Basionym: Danthonia clelandii Vickery, Contrib. New South Wales Nat. Herb. 1: 297 
(1950). Rytidosperma clelandii (Vickery) Connor and Edgar, New Zealand J. Bot. 17: 
332 (1979) 
Type: South Australia: Upper Waterfall Gully, Cleland H. 262, 30 Oct 1926 (NSW 
1126, holo!; JBC iso). 
Joycea lepidopoda (Walsh) Linder, comb. nov. 
Basionym: Danthonia lepidopoda Walsh, Muelleria 7: 384 (1991). 
Type: Victoria: 'Bullens Land' Courtneys Road, north of Ash Reserve, Walsh 1709 
(MEL holo!; NSW, BRl iso). 
Thonandia Linder, gen. nov., a Rytidospermate Steud. lemmatis indumento disperso 
breveque quit distincto a pilis superis longisque; et a Notodanthonia Zotov callo 
parva recedit. 
Type: Thonandia longifolium (R. Br.) Linder 
Caespitose or rhizomatous grasses with intra- or extravaginal innovations and 
truncate, two-awned prophylls, plants villous or glabrous. Inflorescence an open 
panicle. Spikelets 6—16 mm long and with 2—7 bisexual florets; disarticulating between 
the florets, callus blunt, villous, approximately as long as rhachilla internode. Glumes 
equal, 3-7-nerved, glabrous to scaberulous along the veins, taller than the floret 
packet, acute. Lemmas 9-veined, bilobed, with the 3 central veins fusing at the base 
of the geniculate awn, and the 3 lateral veins entering the acuminate lemma lobes; 
lemma indumentum of short hairs covering the abaxial lemma surface, with an 
abrupt transition to a row of long, tufted hairs shorter to longer than the lemma 
lobes; lemma lobes at least as long as lemma body; lemma lobe setae shorter than 
the lemma lobes. Awn column flat, cork-screwed, shorter than the lemma lobes; limb 
long and acute. Paleas longer than the lemma sinus, linear, truncate to bilobed, 
keeled, glabrous or villous between the keels and usually with hair-tufts on the 
palea margins. Lodicules 2, cuneate to rhomboid, bristly and with microhairs. Anthers 
3, yellow to orange. Ovary stalked, glabrous; stigmas distant and separated by a 
small double ridge. Caryopsis obovate, embryo half the length of the caryopsis, 
hilum 20-30% of caryopsis length, elliptical to oblong. Chromosome numbers 24,48. 
This new genus differs from Rytidosperma by the short, felt-like indumentum on the 
lemma sharply distinct from the upper row of hairs, and from Notodanthonia by the 
relatively short callus . It contains five species, from Australia and New Zealand. 
Thonandia gracilis (Hook, f.) Linder, comb. nov. 
Basionym: Danthonia gracilis Hook. El. New Zealand 1: 303 t.69B (1853). 
D. semiannularis (Labill.) R. Br. var. gracilis (Hook, f.) Hook, f., Handb. New Zealand FI. 
333 (1864). Notodanthonia gracilis (Hook, f.) Zotov, New Zealand J. Bot. 1: 123 (1963). 
Rytidosperma gracile (Hook, f.) Connor & Edgar, New Zealand J. Bot. 17: 330. 
Type: New Zealand: Rotoiti Lake, Aglionby Plains, Munro 12012 (holo, K). 
Thonandia nigricans (Petrie) Linder, comb.nov. 
Basionym: Danthonia semiannularis (Labill.) R. Br. var. nigricans Petrie, Trans. New 
Zealand Inst. 46: 37 (1914). Danthonia nigricans (Petrie) Calder, J. Linn. Soc. Lond. 
Bot. 51: 8 (1937). Danthonia gracilis Hook. f. var. nigricans (Petrie) Zotov, Trans. Royal 
