Tobe & Raven, Embryology of Onagraceae (Myrtales) 
679 
nucellus, suggest close resemblance between Onagraceae and Lythraceae (for the presence 
of multi-celled ovule archesporium and starch grains in Lythraceae, see Hubert, 1896; 
Joshi & Venkateswarlu 1935a, b, 1936; Venkateswarlu 1937). Third, the combination of 
a crystaliferous endotesta and a fibrous exotegmen indicates relationships of Onagraceae 
with Lythraceae and Sonneratiaceae, but not with other families including Myrtaceae 
and Melastomataceae. Tl-ius we now would conclude that embryological features do 
support the traditional view that Onagraceae are directly related to Lythraceae. We shall 
Figs 23-29. Transverse (23 and 24) and longitudinal sections (25-29) of seeds showing seed and seed 
coat structure in various genera. 23-27. Mature seeds. 28-29. Immature seeds. 23-24. Seed and seed 
coat of Fuchsia jimenezii. 25. Seed coat of Circaea cordata. 26-27. Seed and seed coat of Gongylocaqjus 
rubrkauUs. 28-29. Seed and .seed coat of Xyloimgra arborea. Note tliat the seed coats of Fuchsia and 
dream are thick and have mesotesta (mts), whereas those of Gongylocarpus and Xylonagra are thin 
and lack mesotesta. Additional abbreviations: ce, cellular endospemi; cot, cotyledon; em, embryo; 
entg, endotegmen; ents, endotesta; extg, exotegmen; exts, exotegmen; sc, seed coat. 
