Douglas, Inflorescence development in Carnarvonia 
765 
apparent as the transverse furrow forms (Figs 52-54). The filament is relatively short 
before zonal growth beneath and between each tepal-stamen complex begins (Fig. 46). 
The distal portion of the filament remams free (Fig. 50). The young anthers are 
approximately equal in size by the time zonal growth commences (Fig. 53). The adaxial 
side of the filament remains distinct on the fused tepal-filament interface (Fig. 54). 
Trichomes develop on the tepal-filament interface after trichome initiation on the 
adaxial side of each tepal (Figs 48,54). The trichomes continue to differentiate, obscuring 
the epitepalous condition of the filament (Fig. 50). At anthesis, the anthers dehisce via 
longitudinal lines along the transverse furrow (Figs 60, 65). 
The carpel primordium enlarges and the cleft reaches the base (epeltate — Fig. 55). 
The orientation of the carpels on the two- and three-flowered subunits appear variable. 
However, as is illustrated in Fig. 56, a three-flowered subunit, the carpel in the axil of 
the a-phyll is aligned with the cleft facing 180° away from the subtending a-phyll; 
the carpel cleft in the flower in the axil of the p-phyll faces away from the (i-phyll; 
Figs 62-65. Fig. 62. Dissected ovary showing the initiation of two ovule primordia from lateral 
positions within the loculus. Fig. 63. Dissected ovary sliowing the initiation and development of the 
outer and inner integuments (oi and ii respectively). The ovules are beginning to arch downwards 
into the loculus. Fig. 64. Mature ovule is hemitropous. Fig. 65. Flower at anthesis showing the lateral 
dehiscene of the antherss and the erect carpel. &ale bars: 62-64 = 50 pm; 65 = 500 pm. 
