Crisp & Chandler, Paraphyletic species 
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Appendix 1. Postulated autapomorphies in species and subspecies from genera in Proteaceae and 
Fabaceae for which phylogenetic information is available. The third column lists taxa whose 
taxonomic segregation might have rendered a taxon without an autapomorphy paraphyletic. 
(Sources: Pate et al. 1989; Crisp 1990; Crisp 1991; Taylor & Crisp 1992; Weston & Crisp 1994; Crisp 
1995a; Crisp 1995b, as well as original observations.) 
Taxon 
Autapomorphies Segregants 
Fabaceae: Mirbelieae 
Brachysema 
bracteolosum 
Enlarged sheathing bract 
celsianum 
Short wings 
lati folium 
Calyx lobes attenuate, almost valvate 
melanopetalum 
Petals purple-black 
minor 
None modestum 
modestum 
Stolon-like inflorescences; creamy pink petals 
papilio 
Crescentic leaf with pungent mucro 
praemorsum 
None? (pale leaf margins are not fixed) papilio 
sericeum 
None melanopetalum 
subcordatum 
Deep pink petals 
Chorizema 
aciculare 
Short shoots 
subsp. aciculare 
None subsp, laxum 
subsp. laxum 
Leaves not tightly revolute; petals creamy 
carinatum 
None spathulatum 
circinale 
Leaves circinate 
cordatum 
Pod distinctly stipitate 
cytisoides 
Bracteoles and calyx-lobes longer than calyx-tube 
dicksonii 
Orange-red petals; stigma with a tuft of hairs 
diversifolium 
Leaves herbaceous; keel obtuse or rounded 
genistoides 
glycinifolium 
Leaves shed at flowering 
Leaves heteromorphic, upper ones longer 
and narrower 
humile 
Leaves obovate to obcordate 
ilicifolium 
Leaf triangular with elongate apex 
nanum 
Plant and parts diminutive 
nervosum 
Leaf apex deflexed 
obtusifolium 
Blunt keel 
parvlflorum 
Racemes condensed 
racemosum 
Leaf margins revolute 
reticulatum 
Long erect stems; ovate leaves 
retrorsum 
Leaf prickles retrorse; style straight 
