74 
ERIK A : SON STENSIO 
In the hyoid arch we find in W.sinuosa the two elements characteristic of Coelacanthids 
well developed and apparently in their original position in relation to each other. Of 
these the ossification in the ceratohyal (City-, text fig. 27; PI. 4, fig. 2; PI. 7, fig. 3 ) is 
unusually long and powerful, and, as usual, slightly arched in such a way that it is 
convex on the ventral side and concave on the dorsal one. The ends, which in their 
present condition are truncated, have been furnished with cartilaginous epiphyses. The 
lower margin has a broadly rounded projection somewhat behind the middle. The posterior 
end of the bone reaches a good distance behind the pterygoid; the opposite end extends 
almost as far forward as the anterior limb of the latter bone. 
The second ossified element of the hyoid arch (Eph, text fig. 27), which is usually 
called the stylohyal in the current terminology, I have here interpreted as the epihyal. 
It is short and fairly thick and truncated at the ends. In its interior it was cartilaginous 
— the ossification forms only a thin perichondral covering — and this cartilage was 
undoubtedly continuous with the epiphyses of the bone, which were also cartilaginous. 
The epihyal ossification is widest at 
its dorsal end, from which it narrows 
fairly gently in ventral direction, 
increasing again in width in the 
neighbourhood of the ventral end. 
Its narrowest part is consequently 
situated near the latter end. Its 
dorsal end lies close to the basis 
cranii, postero-medially of the ptery¬ 
goid bone, but in such a relation 
to the latter bone that, as has been 
pointed out above, it cannot have 
been connected with the metaptery- 
Text fig. 27. Wimania sinuosa, n. sp. 
The two ossifications of the hyoid arch. From P. 257. */« 
Eph, epihyal ossification; Chy , ceratohyal ossification. 
goid. It must thus have joined the neurocranium, but it is impossible to decide whether 
this took place directly or whether another cartilaginous dorsal element corresponding to 
a pharyngobranchial was developed.*) As is clearly shown by this exposition, the 
skeleton of the hyoid arch in the Coelacanthids thus presents striking resemblances 
with that of the Dipnoi, both by the reduction in its dorsal part and with regard to the 
shape of the ceratohyal ossification (cf. Gaupp, 1904, pp. 920—*g 3 i; van Wijhe, 1882, 
pp. 295—297; Ruge, 1897, pp. 257—-284; Furbringer, 1904, pp. 490—495). 
In the species in question the operculum is particularly high and broad. Its contours 
are, however, to a great extent badly preserved, so that the shape cannot be established 
with certainty. Similarly it is impossible to decide whether there was sculpture or not. 
There are five pairs of gill-arches, which are ossified in two segments. The ventral 
of these segments, which I take to be a ceratobranchial (text figs. 28 A, B; PI. 4, fig. 1; 
PI. 7, fig. 3 ) is particularly long and arched in such a way that it has the convexity 
backwards and downwards and the concavity upwards and forwards. Its ventral end is 
q On the left side of the head in question (P. 257) there is a detached bone (y, PI. 4, fig. 2) laterally of 
the posterior end of the ceratohyal ossification. This bone probably belongs to the pectoral girdle or the endo- 
skeleton of the pectoral fin. 
