262 
AMERICAN JOURNAL OF BOTANY 
[Vol. io, 
when selfed produced 46 hermaphrodites of varying degrees of fertility, 
and 2 males which were completely sterile. As will be shown later, the 
pistillate strawberry plants appear to be heterozygous for sex, and therefore 
the female grandparent F. virginiana would be of the constitution FM and 
when crossed with 778 ( HH ) should give 1 female ( FH ) to 1 hermaphrodite 
( HM ). HM was the supposed constitution of the hermaphrodite selfed in 
lot 57/16. The expectation from it when selfed would be 1 HH : 2 HM : 
1 MM. There were produced 46 hermaphrodites and 2 males, one of which 
was very weak. The hermaphrodites varied in fertility. Five were com¬ 
pletely fertile in all flowers; 26 set all but quaternary flowers; 14 were 
fertile in primary and secondary flowers while the tertiaries were either 
sterile or produced nubbins and the quaternaries were sterile; 1 produced 
only nubbins and was practically a male. Although it is difficult to classify 
each of these with respect to its genetic constitution, it seems apparent 
that all the expected classes are represented; the HH being those completely 
or nearly completely fertile, the MM those completely sterile, and the 
HM group those showing intermediate degrees of fertility. These results 
are significant of the constitution of the wild female, as they show conclu¬ 
sively that the tendency toward sterility of a dioecious species may be 
transmitted by the wild female, which is fertile, to hermaphroditic progeny; 
whereas its female progeny are always completely fertile. A probable 
explanation for the variations in fertility of the HM individuals will be 
discussed later. 
Lot 15/16, Glenville X self, although conforming to expectation as far 
as somatic characters are concerned, was strikingly different as to fertility 
from the other selfed hermaphrodites. This variety was mentioned in a 
previous paper (7) as being practically sterile when grown under field 
conditions although it produced blossoms profusely. When grown in a 
bench in the greenhouse it exhibited the same sterility in the first crop of 
blossoms, but when potted and kept in the house until a second crop of 
blossoms was produced it exhibited a fairly high degree of fertility. It is 
evident that environment plays an important part in the fertility of certain 
clones. This variety (15/16) when selfed produced 55 seedlings, 27 of 
which were completely sterile under field conditions while 28 produced 
some fruit. Of these, 11 exhibited a fairly high degree of fertility under 
field conditions, while the remainder set only an occasional flower and the 
berries produced were often nubbins. One seedling, which produced an 
occasional berry when grown in the greenhouse, was self pollinated (54/16). 
It produced 5 seedlings all of which were sterile to a high degree, setting 
only an occasional achene on some of the primary flowers. It appears from 
the results obtained that Glenville carries an hermaphrodite factor which 
is fertile and another which is practically sterile. Under certain cultural 
conditions, the first of these factors is dominant and fruit results, while 
under other conditions the other is dominant and the plant is sterile. 
