May, 1923] VALLEAU—-INHERITANCE IN THE STRAWBERRY 
265 
Pistillate X Hermaphrodite 
Table 3. Results of Crossing Pistillate with Perfect-flowered Varieties of Strawberry 
Lot 
No. 
Varieties Crossed 
Se? 
Hermaph¬ 
rodite 
i of Fi Seedlin 
Pistillate 
igs 
Staminate 
28/16. 
(Fi of Crescent X ?) 
21 
X Minn. no. 3 
16 
40/16. 
Enormous X Dunlap* 
Bederwood X Glenville* (7 plants) 
14 
11 
40/16. 
Enormous X Dunlap 
5 
7 
40/16. 
Enormous X Dunlap* 
5/15 67-2 9 X Glenville* 
13 
14 
66/16. 
Columbia X 5/15 70-4 
26 
27 
► 5 /i 5 . 
F. virginiana X 778 
18 
18 
33 /i 6 . 
5/15 62-5 X Glenville 
27 
37 
45 /i 6 . 
5/15 64-2 X Glenville 
32 
28 
49/16. 
Bederwood X Glenville 
26 
28 
65/16. 
Columbus X Glenville 
9 
10 
56/16. 
5/15 X Glenville 
8 
5 
Total. 
199 
201 
* Plants mixed by acident. 
If we accept the chromosome theory of inheritance as applied to sex in 
the strawberry, we must conclude that the females are heterozygous for 
the determiners H or M and F, depending on whether cultivated varieties 
or wild pistillates are concerned, in view of the fact that the male plants 
and the hermaphrodites have both been shown to be homozygous for 
sex determiners. 
The results of crosses in which pistillates were used as the female parent 
and various hermaphrodites as the male parent are given in table 3. There 
is a definite 1 : 1 segregation of the 400 seedlings into 199 hermaphrodites 
and 201 pistillates. 
In four crosses of pistillate X hermaphrodite made by Mr. Charles 
Haralson and recorded by the writer, the results shown in table 4 were 
obtained. 
Table 4. Results of Crossing Pistillate with Perfect-flowered Varieties of Strawberry 
Pistillate 
Hermaphrodite 
Seedling 1020 9 X Progressive. 
. 31 
4 i 
Seedling 1020 9 X Seedling 907. 
. 16 
9 
Meteor 9 X Seedling 907. 
. 28 
22 
Productive 9 X Seedling 1017. 
. 4 
5 
Total. 
. 79 
77 
These results leave little doubt that the pistillate plants of Fragaria are 
heterozygous with respect to the sex-determining factors. The results of 
the writer in crossing pistillates by hermaphrodites are in keeping with the 
