AMERICAN 
JOURNAL OF BOTANY 
DEVOTED TO ALL BRANCHES OF 
BOTANICAL SCIENCE 
ESTABLISHED 1914 
EDITED BY A COMMITTEE OF THE 
BOTANICAL SOCIETY OF AMERICA 
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EDITORIAL COMMITTEE 
C. E. Allen, Editor-in-Chief, 
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University of Wisconsin 
William Crocker, 
The Thompson Institute 
For Plant Research , Inc. 
C. Stuart Gager, Business Manager 
Brooklyn Botanic Garden 
R. A. Harper, 
Columbia University 
H. H. Bartlett, 
University of Michigan 
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Arthur IK Graves, 
Brooklyn Botanic Garden 
I. F. Lewis, 
University of Virginia 
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The journal is published monthly, except during August and September. Subscription 
price, $6.00 a year. Postage to Canada, 20 cents; to other foreign countries (except Mexico and 
Cuba) 40 cents a volume on annual subscriptions. Single copies 75 cents. Back numbers, 75 cents. 
Back volumes 3 and 5, $8.00; other back volumes, $7.00, post free. 
Donald Reddick, Cornell University 
(Representing American Phytopathological Society) 
The pages of the Journal are open to members of the Botanical Society of America, or to 
candidates approved for membership. 
Manuscript offered for publication should be typewritten doubled-spaced, and should in al 
cases be submitted to the Editor-in-Chief. 
Papers are limited to 20 pages in length, including plates, except that additional pages may 
be arranged for at cost rate to authors (approximately $4.60 per page). 
Illustrations and tabular matter together should, in general, be limited to approximately 
ten percent of the total number of pages. Authors may be charged at cost rates for illustrations 
and tabular matter in excess of that amount (approximately $8.40 per page for tabular matter; 
half-tones $8.30 per page, $5.80 per half page; zinc etchings $5.50 per page, $3.90 per half page). 
ilf-tones, and $2.50 each for zinc etchings. 
For smaller figures the minimum charge is $3.75 each for hal 
Proofs should be corrected immediately on receipt, and returned to American Journal of 
Botany, Brooklyn Botanic Garden, Brooklyn, New York. 
Separates should be ordered from the printer When proof is returned, 
cover will be supplied free; cover and additional copies at cost. 
Remittances should be made payable to American Journal of Botany. 
Fifty copies without 
Claims for missing numbers should be made within 30 days following their date of mailing. 
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The publishers will supply missing numbers free onjy when they have been lost in the mails 
Correspondence concerning editorial matters should be addressed to Prof. G. E. Allen, 
University of Wisconsin, Madison, Wisconsin. 
Business correspondence, including notice of change of address, and directions concerning 
reprints, should be addressed to American Journal of Botany, Brooklyn Botanic Garden, Brooklyn, 
N. Y„ or Lancaster Press, Incorporated, Lancaster, Pa. 
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