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Monocotyledons with Petciloid Flowers — OrcJndacecz . 
STAMEN consolidated with a continuation of the pistil above the sessile stigma forming a “column” opposite to 
the lip. POLLEN various, powdery or coherent in waxy or granular masses : when coherent (forming pollinia) with 
or without a disk-like appendage (gland) united to the pollinia by a slender pedicel ( caudicle ). Upon characters 
derived from the structure of the column and of the pollen Orchidacese are divided into tribes, of which the more 
important are:— 
* ANTHER ONE. 
1. Malaxide^E. — Pollen coherent in waxy masses, unappendaged ; as in the British genera Liparis and Malaxis 
and the Tropical Asiatic genus Dendrobium cultivated in plant-houses. 
2. Epidendre^e. —Pollen coherent, provided with a caudicle but destitute of a distinct gland. There is no 
European representative of this group. Gattleyct and Epidendrum of plant-houses are examples. 
3. VANDEAL —Pollen coherent, provided with a distinct gland, to which it is usually connected by a caudicle. 
Nearly all tropical; none European. Examples : Gatcisetum , Vanda, Oncidium, Saccolabium of plant-houses. 
4. Opi-IRYDEJE.—A nther erect. Pollen consisting of numerous minute granular segments united into 2 pollinia 
by concealed elastic threads, which are continued in each pollen-mass into a caudicle terminating in a gland, or (as 
in Pyramidal Orchis) both caudicles united to the same gland. To this Tribe belong the British genera Orchis , 
Habenaria (including the Butterfly and Frog Orchis) and Ophrys (including the Fly and Spider Orchis). 
5. Neotte^e. —Anther erect or resting obliquely upon the back of the column. Pollen granular, loosely coherent, 
destitute of caudicle or gland in British genera, but with the anterior end of the friable pollinia in immediate contact 
with a secreting organ which emits viscid fluid on irritation. Examples : Bird’s-nest and Tway-blade {Listera). 
* ANTHERS TWO. 
6. Cypripedie^E. —Represented by a single extremely rare indigenous species in Britain, Lady’s Slipper 
(Cypripedium ), so called from its hollow, slipper-shaped lip. 
Ovary inferior, 1-celled, with indefinite ovules upon 3 parietal placentas. STIGMA usually a concave viscid disk 
under the column. 
