DE CANDOLLE ON THE JTIGLANDE J2. 
871 
“ reason for separating it from Mastworts, except its resinous 
“juices.” He nowhere goes the length of saying the walnut has 
either a many-celled fruit or an acorn-cup. It is by no means clear 
to us that Juglandese could have been better placed than in the 
‘ Vegetable Kingdom.’ The divided stigma of the Order appears to 
us as important an item in estimating the relationship between wal¬ 
nuts and oats as is that of the crumpled cotyledons of some oaks, 
advanced by Hr. Lindley. This indicates the essentially com¬ 
pound character of the ovary in the walnuts, while Corylacese, on the 
other hand, with this compound character yet a stage more fully ex¬ 
pressed in the development of dissepiments in the ovary, are never¬ 
theless almost invariably one-celled in fruit. It does not appear 
whether the accidental occurrence of two or more-celled Juglandese 
has ever been observed. The relationship of the group to Anacar- 
diacese adopted from Jussieu by the elder He Candolle, the author of 
this Memoir, as a dutiful grandson, appears more willing to favour. 
His suggestion of an analogy with Myricacese, a small and neglected 
order, is interesting. 
The Memoir is divided thus: Chapter I. Generality. Bistri- 
bution Geograpliique.—II. Organes de la Vegetation.—III. Organes 
de la fructification.—IV. Especes et Varietes Nouvelles.—V. Bivi¬ 
sion de la Eamille et ses Affinities.—VI. Juglandees Eossiles. 
The most important observations appear to be those upon the 
organogeny of the flowers, and the common types to which the male 
and female may be respectively referred. In the walnut, the male 
flowers are arranged in lateral catkins, each flower being subtended 
by a small bracteole, along which bracteole the base of the 6-lobed 
perianth is aduate. This perianth contains a variable number of 
stamens, in two or more rows, the outer set alternating with its 
divisions. The two outer perianth-lobes are antero-lateral, denoted 
by the figures 1 and 2 in the adjoining- 
diagram, b representing the subtending 
bracteole. The next following lobes are 
anterior, 3 ; and posterior, 4; the inner- 
2. most two are postero-lateral, 5 and 6. In 
Ca?ya, M. He Candolle describes the 
perianth of the staminate flower as 3-lobed, 
one of the lobes being anterior and exterior. 
This is the bractlet. The perianth proper, we should say, is reduced 
to 2 lobes, answering to the lobes 1 and 2 in the walnut. These, 
however, are adnate with b , and form a single envelope. Thus, in 
Cary a we have the flower of Juglans with the perianth lobes, 8, 4, 
5, and 6, undeveloped. 
With regard to the female flowers of these genera, in the walnut 
the six lobes of the perianth and the bracteole are united together 
with the ovary. The bract and perianth-lobes, 1 and 2, constitute 
the outer, the four remaining perianth-lobes the inner perianth. In 
the pistillate flower of Cary a the bract and three of the lobes unite 
N. H. R.—1863. 2 C 
4. 
5 . 6 . 
1 . 
3 . 
b. 
