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the same wild mutation elsewhere. The flowers have five spurs 
and this type is now known as pelovia nectaria. There is another 
type, pelovia anectavia , in which the flowers are actinomorphic, but 
without spurs. Another race studied by de Vries, which he called 
hemipeloria , produces usually a single (terminal) peloric flower on 
each plant. Such a flower may not appear in some years, or 
occasionally more than one may appear, and this seems to be 
controlled to some extent by the environment, but the capacity for 
producing a peloric flower is evidently inherited through the 
seed. In breeding from such a half-race de Vries finally obtained 
mutations to the full peloric condition, in the fifth and sixth 
generations of culture. This mutation occurred with a frequency 
of about 1%, and although highly sterile it probably breeds true. 
Whether there are any plants which never produce a peloric flower 
is uncertain. 
A similar peloric condition (Digitalis purpurea monstrosa ) has 
long been known in foxgloves, in which a single, erect terminal 
peloric flower occurs on the plant, and opens before the other 
flowers, which then follow in acropetal succession. The peloric 
flower frequently also shows an increase in the number of parts. 
This condition is found by Keeble, Pellew and Jones (1910) to be 
inherited as a simple Mendelian recessive. The peloric flowers are 
perfectly fertile with their own pollen, the inheritance being the 
same through peloric or normal flowers of the plant. 
In Antirrhinum majus t as in many other plants with 
zygomorphic flowers, sporadic peloric flowers occasionally occur. 
There is also a completely peloric race, which Baur (1911) used in 
his extensive crosses. He concluded that zygomorphy depended 
upon the presence of two factors P and E. In the absence of E 
the plant will have only peloric flowers, but plants having E without 
P (EEpp) produce both peloric and normal flowers. This is an 
interesting case of somatic segregation in the presence of one pair 
of factors, and presumably there would be no difference in 
inheritance from the two types of flower on the same plant. If 
such an EEpp individual is crossed with a fully peloric race 
(eePP), the (EePp) bears only normal flowers, while the P 2 gives 
(1) normal-flowered plants, (2) plants with both normal and peloric 
flowers, (3) plants with only peloric flowers, in the ratio 9:3:4. 
The numbers actually obtained were 70:13: 45, and the departure 
from expectation (72 : 24 : 32) is explained by the fact that badly 
nourished plants belonging in class (2) produce only peloric flowers 
