ANOMACANTITUS : A NEW GENUS OE ACANTHACE2E 1G3 
absent. The two posterior stamens are complete and two-celled, 
while the anterior ones are one-celled (cf. Brillantaisia). Owing to 
the curvature of the filaments the stamens lie, almost in contact, 
obliquely across the corolla tube. The anthers are long and narrow, 
Avith sharply hooked tips and are furnished at the base of each ceil 
with a glandular cushion-like mass. Both anthers and filaments bear 
numerous capitate glandular hairs. The fruit is remarkably large 
for the family and is drupaceous. Within the fleshy exocarp are two 
large pyrenes, subglobose but flattened on the side of contact. In 
the type-specimen the outer fleshy la} r er is incomplete and of un¬ 
certain thickness. The pyrenes are woody and hard, much corrugated 
on the outer surface and lined within by a dense layer of brown felt. 
The seeds are large and without a hard coat. The placentae are large 
and project as sharp ridges into the cavity of the pyrenes. In the 
young ovary the septum is incomplete, but in the ripe fruit structures 
can be seen which appear to be the result of subsequent completion 
of the septum. 
The leaves are without cystoliths. The epidermal cells are sinuate 
in outline, while the stomata are of normal Acanthaceous type. 
Typical subsidiary cells are present. 
The minute structure of the stem also shows many points of 
interest. In transverse section it is roughly quadrangular, with rather 
prominent corners and concave sides. The stem is hollow and the cen¬ 
tral cavity occupies about f of the total diameter. Broadly the stem is 
a thin-walled tube strengthened at the cardinal points of its circum¬ 
ference by 4 massive square-faced ribs. Additional ribs are subse- 
sequently formed. Beneath the epidermis is a well-marked hypoderm 
consisting of a single layer of stone cells, forming a complete ring 
below the surface of the stem. Within this are two further narrow 
rings of tissue. The outer is dark in colour and almost structureless. 
The inner is very open, with numerous scattered fibrous cells. These 
two layers seem to represent, respectively, the endodermis and the 
pericycle. The innermost tissue in the stem is a broad ring of 
centripetal secondary xylem, the elements of which are more or less 
rounded in section. On the inside, the edge of the stem-cavity is 
sharply defined and all trace of primary xylem is lost. Each of the 
4 main stem-ribs consists of a 4-sided mass of centrifugal secondary 
xylem. The elements of this wood are square in section and regularly 
arranged. In each rib there are from 8 to 10 very large vessels. 
The outer extremities of each xylem mass are occupied by particu¬ 
larly large vessels, while those inside are smaller. Medullary rays, 
one cell broad, are present in the centrifugal xylem, but are absent 
from the centripetal ring. Phloem is poorly developed. A certain 
amount appears to occur between the main centrifugal xylem masses. 
The centrifugal xylem appears to be of comparatively late incep¬ 
tion in response to the assumption, by the young plant, of the 
climbing habit. In function it gives necessary strength to the stem 
and provides the large and conspicuous vessels, and is the result of 
the production of wood elements outwardly by the cambium. During 
this process the outer stem-tissues become much compressed and the 
phloem becomes unrecognisable. Where this crushing effect is 
