Notes . 
156 
The facts detailed above make it clear that we are concerned with a member of 
the Peronosporineae which cannot readily be referred to any of the existing genera, 
though certain of the facts detailed above indicate a close 
Fig. 3. Portion ot mycelium 
from artificial culture show¬ 
ing conidia, c , and oogonia, 0. 
x 140. 
relation to the genus Phytophthora . The amphigynous 
antheridium has hitherto been found only in certain 
species of this genus,andLafiferty and Pethybridge (loc.cit.) 
in a recent communication have given a list of fourteen 
species in which this type of antheridium has been found, 
either alone or associated with the paragynous form, 
whereas in five species the latter has alone been recorded. 
If the character of the antheridium be regarded as 
indicating some affinity to Phytophthora , the sacculate 
oogonia mark it off from the species of this genus, 
though somewhat similar forms have been described in 
members of both the Ancylistineae and Saprolegnineae. 
Moreover, the conidiophores are quite unlike those of 
Phytophthora or indeed of any other genus of the family, 
and the same is true of the large echinulate conidia. 
• 
If the conidial fructification be considered alone, 
it might be possible to associate it very closely with the 
genus Muratella , described by Bainier and Sartory, 1 
which these authors are inclined to consider a member 
of the Mortierellaceae. In both, the conidia are echinu¬ 
late and borne on globular enlargements of the conidio¬ 
phores, though in Muratella these are usually terminal. 
The uncertain position of Muratella, however, and the 
fact that in the fungus under consideration the sexual 
mode of reproduction is known, has inclined the authors 
to place it in a new genus of the Peronosporineae, based 
on the characters of the conidiophores and the sacculate 
oogonia, under the name of Trachysphaera fructigena. 
Trachysphaera , nov. gen. (rpcixvs = rough, o-paipa = 
globe). 
Conidiophorum simplex vel ramosum vesiculis ter- 
minalibus vel intercalaribus conidia pedicellata gerentibus 
praeditum. 
Antheridia amphigynosa. Oogonia pyriformia e 
bulloso alte tuberculosa, oosporas episporio tenui vestitas 
liberas continentia. 
Trachysphaera fructigena, nov. sp. 
Mycelium intercellulare plerumque continuum non-septatum e hyphis crassius- 
culis compositum; haustoriis nullis. 
1 Bainier, G., et A. Sartory: Etude morphologique et biologique du Muratella elegans. Bull. 
Soc. Mycol. France, xxix, 1913, pp. 129-36. 
