427 
Physiology of the Genus Eidamia. 
borne terminally on lateral branches. The differences lie in the form of the 
conidiophore, the appearance of the conidia in groups in the case of E. viri- 
descens , and the fact that the macrospores are hyaline instead of brown in 
both fungi. It can be shown that these differences break down when 
growth takes place on different media. 
The conidiophore of E. acremonioides is typically of the Aspergillus 
form, the globular apical portion bearing sterigmata with spores in chains. 
Under certain cultural conditions the sterigmata are borne, either singly or 
in small lateral or terminal groups, on undifferentiated branch hyphae. 
When Fig. 3 (E. acremonioides ) is compared with those illustrating the 
conidiophores of E. viridescens the similarity in position of the sterigmata 
and their size and shape is very striking. In the case of E. catenulata the 
conidiophores are usually not apically swollen, but approximations to the 
types found in E. acremonioides do occur (Fig. 10). 
In Eidamia the conidia are typically borne in chains, but, even in the 
case of E. acremonioides , a disposition in groups at the apices of the sterig¬ 
mata is not uncommon. In E. viridescens grouping is the rule and catenu- 
late conidia are rare. From the occurrence of intermediate stages (Fig. 12, 
E. catenulata) a distinction between chains and groups of spores as a criterion 
of specific value is of little importance in the genus Eidamia. The conidia 
in the two new fungi are coloured in contrast to the colourless state of those 
of E. acremonioides , but this difference is not of generic value. 
The macrospores of E. acremonioides are relatively larger than those of 
the other two species. They are usually brown in colour as contrasted with 
the lack of colour in E. catenulata and E. viridescens , though occasionally 
hyaline spores are produced (potato mush agar at 30° C. or potato slab at 
25°C.). Therefore the two new fungi cannot be excluded from the genus 
owing to the hyaline character of the macrospores. From the foregoing it 
will be seen that the differences between the three fungi are not of sufficient 
importance to prevent their inclusion in one genus. 
The bulbils found in E. acremonioides (Papidaspora aspergilliformis) 
and in Helicosporangium parasiticum by Eidam (6) are regarded by 
Bainier (1) as perithecia containing an ascogonium capable of developing to 
form atypical perithecium, with ascospores set at liberty through an ostiole. 
Normally the'development appears to be arrested when the bulbils contain 
central cells surrounded by a sheath. After resting, such a bulbil is capable 
of growth by the production of vegetative filaments. Moreau ( 12 ) has 
worked out the cytology of these structures, and his conclusion supports 
Eidam and Bainier in the view that the bulbils are perithecia arrested in 
their development, not abortive, but capable of proceeding with the normal 
formation of a perithecium under certain conditions. No bulbils were found 
in E. acremonioides on the media used in this investigation, nor was their 
presence detected in E. catenulata and E. viridescens. 
