24 
ROBERT S. HILL 
Both specimens yielded well preserved cu¬ 
ticle, although the stomatiferous surface of 
Banksia adunca could not be cleaned completely 
despite a number of attempts. It is possible that 
other methods may be successful in cleaning this 
cuticle, but given the small amount available it 
was considered that the risk of damaging the 
cuticle was too great to persevere. 
Both species clearly belong to the tribe Bank- 
sieae of the Proteaceae. They contain a combi¬ 
nation of characters listed by Cookson & Duigan 
(1950) and Hill & Christophel (1988) as being 
typical of the tribe (stomata with paired subsid¬ 
iary cells parallel to the pore (brachyparacytic), 
serrations with prominent veins ending in the 
apices, and distinctive trichome types). Like 
other fossil leaves of this type, however, they 
cannot be placed with certainty in either Bank¬ 
sia or Dryandra , and so must be transferred to 
the form genus Banksieaephyllum. 
TAXONOMIC DESCRIPTIONS 
Family Proteaceae 
Subfamily Grevillioideae 
Tribe Banksieae 
Genus Banksieaephyllum Cookson & Duigan 
Banksieaephyllum urniforme (Deane) Hill, 
comb. nov. 
Figs 1A-D, 2A 
Dryandra urniformis Deane 1925: 495, pi. 62, 
fig. 9. 
Holotype. NMV PI5127. 
Emended diagnosis. Leaves bilateral, at least 6 
mm wide, pinnately lobed. Lobes acute and api- 
cally directed; apical and basal sides of lobes 
convex, apical side much shorter than basal side. 
Leaf base and apex unknown. Secondary vena¬ 
tion pattern craspedodromous, with a number of 
secondary veins per lobe. Stomata brachypa¬ 
racytic, superficial. Trichome bases of two types 
abundant on stomatiferous epidermis. Large tri¬ 
chome bases usually associated with one but 
sometimes with two or three epidermal cells; tri¬ 
chome small, unicellular. Small trichome bases 
associated with one epidermal cell; trichome 
basal cell thickly cutinized, cylindrical; apical 
cell(s) not preserved. Trichome bases absent 
from upper epidermis. 
Discussion. This species is represented by only a 
fragment of a single leaf but the cuticle is well- 
preserved. The leaf has characteristic urn- 
shaped pinnae which are similar to those de¬ 
scribed in Banksieaephyllum elongatum Hill & 
Christophel (1988). Hill & Christophel (1988) 
commented on this similarity but considered the 
pinnae of B. urniforme and B. elongatum to be 
consistently of a different shape. The specimen 
of B. urniforme was not studied by Hill & 
Christophel, however, and its cuticular morpho¬ 
logy was therefore not considered. The cuticle of 
B. urniforme is similar to that of B. elongatum 
but differs from it most notably in lacking small, 
simple trichome bases on the upper epider¬ 
mis. 
The remaining fossil species of Banksieae with 
pinnate leaves are much larger in size thanto( r - 
sieaephyllum urniforme , and have very distinct 
cuticular patterns ( Banksieaephyllum cuneaium 
Hill & Christophel, 1988 and B. incisum Black¬ 
burn, 1981), distinctive leaf bases (Banksieae' 
formis decurrens Hill & Christophel, 1988), or 
distinctively rounded pinnae ( Banksieaephyl¬ 
lum pinnatum Cookson & Duigan, 1950). 
Banksieaephyllum elongatum Hill & 
Christophel, 1988 
Banksieaephyllum elongatus Hill & Christophel 
1988: 212-214, figs 24-29. 
Emended diagnosis. Leaves bilateral, at least 60 
mm long and up to 10 mm wide, pinnately 
lobed. Lobes acute and apically directed; apical 
side of lobe usually straight or slightly concave, 
much shorter than basal side; basal side convex. 
Leaf base cuneate, apex unknown. Secondary 7 
venation pattern craspedodromous, with a vari¬ 
able number of secondary veins per lobe. 
Stomata brachyparacytic, superficial. Trichome 
bases of two types abundant on stomatiferous 
epidermis. Large trichome bases usually asso¬ 
ciated with one but sometimes with two or three 
epidermal cells; trichome small, unicellular. 
Small trichome bases associated with one epi¬ 
dermal cell; trichome basal cell thickly cutin¬ 
ized, cylindrical; apical cell(s) not preserved. 
Small, unicellular trichome foot cells scattered 
on upper epidermis; trichomes unknown. 
Discussion. An emended diagnosis of B. elonga¬ 
tion is given here because in the original diag¬ 
nosis the small unicellular trichome bases on the 
upper epidermis were not included. This is one 
