2 
N. W. ARCHBOLD 
Genus Permorthotetes Thomas, 1958 
Type species. Permorthotetes callytharrensis Thomas, 
1958. 
Comments. Permorthotetes was regarded by 
Manankov (1979) as a synonym of Orthotetes 
Fischer de Waldheim, 1829, but I agree with 
Manankov’s (1973) earlier view that these gen¬ 
era are distinct. 
Permorthotetes lindneri Thomas, 1958 
Fig. 1A-C 
Permorthotetes lindneri Thomas 1958: 92, pi. 1, figs 
1 -4, pi. 2, figs 1 -3, pi. 3, figs 1 -6.—Guppy ct al. 1958: 
54.—Thomas in Dickins 1976: 99. 
Orthotctid.—Dickins 1976: 100. 
Material. CPC 28024-28025, portion of a ventral 
valve external mould and portion of a dorsal valve 
external mould from locality TK5A, Scott Bluff, 
southern Canning Basin (collected by W.J. Koop). 
Comments. The nature of the flat ventral valve, 
fine costellae and step-like lamellae permit 
assignment of the material to P. lindneri as 
identified by Thomas (in Dickins 1976). 
Thomas (1958: 92-96) fully described P. 
lindneri and compared it with other Western 
Australian representatives of Permorthotetes. 
Order Productida Sarycheva Sc Sokolskaya, 
1959 
Suborder Chonetidina Muir-Wood, 1955 
Superfamily Chonetacea Bronn, 1862 
Family Anopliidae Muir-Wood, 1962 
Discussion. Since the reviews by Archbold 
(1980, 1981), this family has been investigated 
by Afanas’yeva (1984a, b). She stressed the 
appearance of heterochronous homeomorphs 
and considered it preferable to subdivide the 
family on the basis of dorsal internal features 
rather than on external ornament. The sub¬ 
family Tornquistiinae, possessing low radial 
lines of fused papillae in the dorsal interior, was 
separated by Afanas’yeva (1984a) from the 
Anopliinae, possessing paired radial ridges in 
the dorsal interior. Dorsal features do not 
appear to be a reliable criterion for subfamilial 
classification, however, as it is difficult to place 
in separate subfamilies the type species of Ano - 
plia and Tornquistia, the dorsal interiors of 
which possess similar paired interior radial 
ridges (see Muir-Wood 1962, pi. 5, figs 16, 
21 ). 
Subfamily Anopliinae Muir-Wood, 1962 
Genus Tornquistia Paeckelmann, 1930 
Type species. Leptaena (Chonetes) polita McCoy, 
1852. 
Diagnosis. Anopliinids with short lateral septa 
and prominent accessory septa in dorsal in¬ 
terior. Ventral interior with short, thin median 
septum. Shell exterior smooth. 
Discussion. Tornquistia was reviewed by Arch¬ 
bold (1980, 1981) who substantially restricted 
the Permian palaeogeographical range of the 
genus to Western Australia, Thailand, the Salt 
Range in Pakistan, Eastern Siberia and Arctic 
North America. Many Permian species of the 
genus possess dorsal interior paired radial ridges 
that are less distinct than those of the type 
species (e.g. see Afanas’yeva 1977, Archbold 
1980). 
Tornquistia subquadratus sp. nov. 
Fig. lD-O 
“ Chonetes ” sp. — Dickins 1976: 99. 
Holotype. CPC 28026, an internal mould of a dorsal 
valve. 
Other material and locality. CPC 28026-28034, 2 dor¬ 
sal valve internal moulds, 2 dorsal valve external 
moulds, 4 ventral valve internal moulds and 1 incom¬ 
plete ventral valve external mould as well as numerous 
fragments of moulds; all from locality TK5A, Scott 
Bluff, east side of Lake Blanche, southern Canning 
Basin; collected by W. J. Koop. 
Fig. 1. A-C, Permorthotetes lindneri Thomas. A, B, CPC 28024, incomplete external mould of ventral valve and 
latex cast, x 1.5. C, CPC 28025, latex cast of incomplete external mould of dorsal valve, x 1.5. D-O, Tornquistia 
subquadratus sp. nov. D, E, CPC 28026, holotype, internal mould of dorsal valve and latex cast, x 5.0. F, CPC 
28027, ventral valve internal mould, x 5. G, CPC 28029, dorsal valve internal mould and latex cast, x 5. J, CPC 
28030, ventral valve internal mould, x 5. K, CPC 28031, latex cast of dorsal valve external mould, x 5. L, O, CPC 
28032, internal mould of ventral valve and latex cast of mould, x 5. M, CPC 28033, internal mould of ventral 
valve, x 5. N, CPC 28034, latex cast of ventral valve external mould, x 5. 
