48 
M. J. TYLER 
Fig. 1. Comparison of ilia of extant species of Philoria 
and Kyarranus. A, Philoria frosty B, Kyarranus kun- 
dagungan ; C, K. loveridgei ; D, K. sphagnicolus. Scale 
bar = 5 mm. 
developed. Ventral acetabular expansion (VAE) 
missing. Dorsal acetabular expansion (DAE) 
narrow and incomplete. Dorsal prominence 
well- developed and conspicuous. Dorsal 
protuberance projecting laterally and extremely 
conspicuous. Anterior margin of dorsal promi¬ 
nence extends to position located anterior to 
anterior margin of acetabulum. 
Length of ilium 16.1 mm; DAE/VAE cannot 
be estimated with any degree of confidence. 
Paratvpes. There are 15 paratypes: Gag Site QM 
18160, 18166, SAM P31237; Last Minute Site 
QM FI 8163; Neville’s Garden Site QM 
FI8169-70, R.S.O. Site QM FI8161, 18165; 
Camel Sputum Site QM FI 8162, 18168, 18171, 
Fig. 2. Ilia of Kyarranus borealis sp. nov. A, QM 
FI 8169 (paratype); B, SAM P31238 (paratype); C. 
QM FI8167 (holotype). 
SAM P31238; Upper Site SAM P31239; Outa- 
site Site QM FI8172; Wayne’s Wok Site QM 
FI 8164. 
Variation. Fourteen of the 15 paratypes have an 
incomplete shaft. The complete specimen (SAM 
P31238) measures 8.7 mm compared with 16.1 
mm for the holotype. No significance is attri¬ 
buted to this difference beyond ontogenetic; 
comparable variation in size was noted in Lech- 
riodus intergerivus at Riversleigh Station (Tyler 
1989) and the projected sizes of many of the 
incomplete ilia approximate the holotype. The 
proximal portions of each of the ilia, although 
commonly lacking entire dorsal or ventral acet¬ 
abular expansions, conform to the shape of the 
holotype. 
Comparison with other species. It is evident that 
K. borealis may have been a slightly larger 
species than any of the extant congeners. As de¬ 
termined by plotting the regression line ofilial 
length upon snout to vent length of extant 
species (Fig. 3), the holotype of K. borealis would 
have had a snout to vent length of 37.9 ± 0.67 
mm. This figure compares with upper limits of 
31 mm for K. loveridgei , 30 mm for K. kumta- 
gungan and 37 mm for K. sphagnicolus. 
Etymology /. The adoption of borealis (L.) or 
“northern” refers to the far northern position of 
the type locality compared with the geographic 
distributions of extant species. 
