WESTERN AUSTRALIAN PERMIAN BRACHIOPODS 
63 
fold. The specimen shows distinct dimples and a 
few spine bases but is inadequate for detailed 
comparison with other Western Australian 
species described by Coleman (1957). 
Order Spiriferida Waagen, 1883 
Suborder Spiriferidina Waagen, 1883 
Superfamily Spiriferacea King, 1846 
Family Spiriferidae King, 1846 
Subfamily Neospiriferinae Waterhouse, 
1968 
Genus Fusispirifer Waterhouse, 1966 
Type species. Spirifer nitiensis Diener, 1897. 
Comments. Fusispirifer was reviewed by Arch¬ 
bold & Thomas (1987) who discussed the sub- 
familial placing of the genus and its palaeogeo- 
graphical distribution. An additional species of 
Furispirifer from the Western Australian Per¬ 
mian was described by Archbold & Skwarko 
(1988). 
Fusispirifer sp. 
Fig. 3S, 4A-G, L. 
Material. CPC 19894-19897, three ventral valve in¬ 
ternal moulds and one external mould of a ventral 
valve from BMR locality WB 69, basal One Gum For¬ 
mation. CPC 24506-24509, two natural casts of 
juvenile ventral valve interiors and two incomplete 
external moulds of ventral valves from BMR locality 
7864 0427, basal One Gum Formation. GSWA 
F43851 (formerly FI 1060/1), incomplete internal 
mould of a ventral valve from GSWA locality 94217, 
Billidee Formation. 
Comments. Fusispirifer is abundant in the Per¬ 
mian of Western Australia (Archbold & Thomas 
1987) and the present material indicates the 
presence of the genus in the Wooramel assemb¬ 
lages. The species is characterised by coarse cos¬ 
tae and lateral plications that are variable but 
usually well developed on juvenile specimens 
but are poorly developed on the flanks of mature 
specimens. The Callytharra Formation species 
F. carnarvonensis possesses lower and usually 
finer ventral valve costae and very subdued lat¬ 
eral flank plications when compared with the 
material to hand. The younger F. byroensis 
(Glauert) (see Archbold & Thomas 1987) is a 
distinct flattish species readily distinguished 
from the Wooramel Group species. 
A distinctive internal feature of the Wooramel 
species is the large, relatively wide ventral 
muscle field recalling that of the large species F. 
wandageensis from the late Early Permian Wan- 
dagee Formation. The Wooramel specimens are 
closest to F. carnarvonensis from the Callytharra 
Formation but are distinct from that species and 
with the collection of additional material, par¬ 
ticularly dorsal valves, will require the recog¬ 
nition of a new species. 
Genus Neospirifer Fredericks, 1924 
Type species. Spirifer fasciger von Keyserling, 1846. 
Neospirifer sp. 
Fig. 4H-J 
Neospirifer sp .—Konecki et al. 1958: 33.—Condon 
1962a: 11.—Condon 1967: 114.—McGann 1976: 
51. 
Neospirifer sp. B.—Archbold & Thomas 1986: 150, 
fig. 14 B, C. 
Material. One natural cast of a ventral valve interior 
and one incomplete external mould of a ventral valve 
(CPC 19898-19899) from BMR locality 7864 0427, 
and one incomplete external mould of a ventral valve 
(CPC 24362) from BMR locality WB 69; all from the 
basal One Gum Formation. 
Comments. As noted by Archbold & Thomas 
(1986: 150), the Wooramel Neospirifer species is 
related to the Western Australian N. hardmani- 
N. plicatus group of Neospirifer and appears to 
represent an intermediate species between the 
Callytharra and Madeline Formation species. 
The new material includes a fragment of a mat¬ 
ure ventral valve (CPC 19899) which recalls 
specimens of an undescribed species of Neospir¬ 
ifer from the Aktastinian High Cliff Sandstone 
of the Perth Basin. 
Order Athyridida Dagys, 1974 
Suborder Athyrididina Boucot, Johnson & 
Staton, 1964 
Superfamily Athyridacea M’Coy, 1844 
Family Athyrididae M’Coy, 1844 
Subfamily Athyridinae M’Coy, 1844 
Genus Cleiothyridina Buckman, 1906 
Type species. Atrypa pectinifera Sowerby, 1840. 
?Cleiothyridina sp. 
Fig. 4K 
Comments. A single specimen (CPC 24510) of 
an incomplete natural cast of a juvenile ventral 
valve interior from BMR locality 7864 0427 
may represent an athyrid such as Cleiothyridina , 
judging from what appear to be traces of vascu- 
