26 
T.A. Darragh 
inner lip with 17-21 well-developed teeth, extending along 
entire lip. Fossula well developed, deeply depressed, bounded 
anteriorly by sharp terminal ridge. Colour pattern present on 
some specimens, of brown ground with darker brown patches. 
Dimensions. 
L 
W 
H 
LT 
CT 
Holotype WAM 89.177 
57 
41 
34 
27 
21 
Paratype WAM 89.437 
55 
38 
30 
18 
18 
Paratype NMV P308704 
51 
36 
30 
20 
19 
Type locality. Quarry 2.5 km north of Hampton microwave 
repeater tower, Roe Plains, Western Australia. AMG Eucla 
1:250,000 sheet CK365465. Roe Calcarenite. 
Type material. Holotype WAM 89.177, collected G.W. Kendrick 
27-30 October 1988; paratype WAM 89.437, collected Sam 
Rowe, January 1989; paratype NMV P308704, collected T. A. 
Darragh, 9 August 1973. 
Time range. Pliocene. 
Occurrence and material. Roe Calcarenite: PL3172 Hampton 
Tower (P308704-5, WAM 69.494, 70.17, six specimens); 
PL3167 1.5 km north of Hampton Tower (P308703, WAM 
80.109, three specimens); PL3166 2.5 km north of Hampton 
Tower (P121293, WAM 89.178, 89.437, 89.637, four specimens). 
Remarks. At first glance, small specimens of this species may 
be confused with the common Austrocypraea amae Fehse and 
Kendrick of the Roe Calcarenite, but they are readily 
distinguished by their smooth fossula. In Austrocypraea, the 
columellar teeth are produced into thin ribs that continue 
across the fossula. This species is most closely related to the 
living species Zoila decipiens (Smith, 1880), Recent, Western 
Australia, from which it differs by having stronger teeth and 
teeth present along the entire columella. It is also globose in 
shape rather than pyriform as in Z. decipiens and lacks the 
prominent protruding spire of that species. The fossula is very 
similar to that of the Z. decipiens. It is not closely similar to the 
fossil species of Zoila known from the Oligocene and Miocene 
of southeast Australia, with the exception of a species known 
from a single broken specimen from the upper Miocene of 
Victoria. Z. campestris bears some resemblence to Zoila 
kendengensis Schilder, 1941 from the Pleistocene Putiangan 
Formation of Java, but that species has relatively prominent 
anterior and posterior canals. 
Etymology. Latin campester, pertaining to a plain. 
Zoila fodinata sp. nov. 
Figures 16K-L, N 
Description. Shell solid, polished, of average size for the genus, 
globose, pyriform, ventral surface rounded. Spire slightly 
protruding beyond last whorl, covered with thick callus. 
Posterior canal short, notched, sides thickened. Anterior canal 
very short, abruptly truncated, deeply incised. Aperture 
sinuous, widened above fossula; outer lip with 25-28 well- 
developed teeth, extending along entire lip; inner lip with 21- 
25 well-developed teeth, extending along entire lip. Fossula 
well developed, subrectangular, concave, bounded on inner 
side by low ridge and anteriorly by sharp terminal ridge; 
terminal ridge rather broad, extending down into aperture as 
sharp ridge forming anterior edge of fossula. 
Dimensions. 
L 
W 
H 
LT 
CT 
Holotype WAM 89.637 
72 
47 
39 
28 
24 
Type locality. Quarry 2.5 km north of Hampton microwave 
repeater tower, Roe Plains, Western Australia. AMG Eucla 
1:250,000 sheet CK365465. Roe Calcarenite. 
Type material. Holotype, WAM89.637, collected G.W. 
Kendrick, 27-30 October 1988. 
Time range. Pliocene 
Occurrence and material. Roe Calcarenite: PL3166 2.5 km 
north of Hampton Tower (one specimen). 
Remarks. This species is most closely similar to the living 
species Zoila venusta, from which it differs by having stronger 
teeth and teeth present along the entire columella. The fossula 
is also shallower but deeper within the aperture than in Z. 
venusta. It is not as globose, being more pyriform. It does not 
seem to be closely similar to the fossil species of Zoila known 
from the Oligocene and Miocene of southeast Australia. 
Etymology. Latin, fodina, a quarry. 
Acknowledgments 
The following persons have assisted me by providing 
information, and the donation or loan of specimens, and are 
gratefully acknowledged: Chris J. Goudey, Lara, Victoria; 
Peter Hunt, Adelaide, South Australia; David T. Dockery, 
Mississippi Bureau of Geology; the late Warren Blow, United 
States National Museum; Wolfgang Grulke, Dorset, England; 
Ben McHenry, South Australian Museum; George Kendrick 
and Helen Gore, Western Australian Museum; Ian Loch, 
Australian Museum; John Cooper, Natural History Museum, 
London; Barry Wilson, Perth, Western Australia; Pierre 
Lozouet, Philippe Bouchet and Virginie Heros, Museum 
national d’Histoire naturelle, Paris; Alan Beu and John Simes, 
GNS Science, Lower Hutt, New Zealand. I am grateful to 
Rodney Start, Museum Victoria and Mark Darragh for 
considerable assistance with photography. I thank Alan Beu 
and Barry Wilson, who provided significant helpful comments 
and suggestions that have considerably improved the 
manuscript. 
References 
Beu, A. G. and Darragh, T. A. 2001. Revision of southern Australian 
Cenozoic fossil Pectinidae (Mollusca, Bivalvia). Proceedings of 
the Royal Society of Victoria 113(1): 1-205. 
Bouchet, P. and Rocroi, J-P. 2005. Classification and nomenclator of 
gastropod families. Malacologia 47(1-2): 1-397. 
Cernohorsky, W. O. 1971. Fossil and Recent Cypraeacea (Mollusca: 
Gastropoda) of New Zealand with descriptions of new species. 
