A GENETICO-PHYSIOLNGICAL STUDY ON THE FORMATION ETC. 
43 
(3) Apiculus of glumes coloured, but leaf-slieatlis colourless. 
(4) Apiculus of glumes only coloured. He doubts of classes 3 and 4 
really exist. Class 1 is the commonest group. Contrary to Indian varieties, 
the colourless stigma with green leaf-sheath is the most common type found 
in Japanese varieties. He observed further, that the colour in the leaf-sheatlis 
and apiculus is due to a colour factor R acting on a chromogen G, and that 
the purple colour of the stigma is due to a further factor P not present in 
the leaf-sheath and apiculus, and that the simultaneous presence of all three 
factors R CP is necessary for the production of any sort in the stigma. Parnell, 
Rangaswami, Agyangar, and Raniah (1917) 1 have shown that in certain Indian 
varieties, the purple lining of the internode is coupled with the purple glumes, 
the purple stigma with the pimple axil (purple-colouring of the epidermis on 
the inside of the sheath) but the purple lining of the internode and the purple 
glume do not co-exist with the purple stigma and the purple axil. 
Among the Japanese varieties studied by the writer, most of those which 
liave a coloured stigma have a purple awn and leaf-sheaths striped, and only 
in certain varieties, they are separated. The distribution of anthocyanin 
in Japanese varieties is shown in the following table. 
Table 18. 
Distribution of anthocyanin in the varieties of Japanese rice. 
Rame of typical 
variety 
Stigma 
Awn 
Palea 
(tip) 
Palea 
Glume 
Lf. sheath 
Lf. blade 
“ Murasaki ” 
Coloured 
Purple 
Purple 
Purple 
Purple 
Purple 
Purple 
“ Tokachi-wase ” 
99 
99 
99 
Red 
Red 
Purple 
striped 
Red 
striped 
“ Haguro ” 
99 
99 
99 
Purple 
Purple 
Red 
strip. 
— 
“ Edowase ” 
99 
99 
99 
— 
— 
99 
— 
“ Bozu-karasu ” 
99 
Awnless 
99 
— 
99 
— 
— 
“ Uyejini-shirazu *’ 
99 
Purple 
Reddish 
brown 
— 
99 
— 
— 
“ Genroku-mochi ” 
99 
99 
— 
Purple 
Purple 
Purple 
striped 
1 
1. Parnell, F. P-, Rangaswami, Agyangar, G. N, and Raniah, K, The Inheritance of 
Characters in Rice I. Memoirs Depart. Agri. India. Bot. Series. 9 :75, 1917. 
