130 
S. IKENO : 
The agreement between the theoretical and the actual numbers is not very 
bad in both cases in view of the small number of plants, except in respect 
to the orange where the number of individuals is much smaller than might 
be expected theoretically ; this may be due possibly to the fact that a 
comparatively large proportion of seeds of this class of zygotes failed to 
germinate. The further behaviour of one of the two reds segregated out in 
F ?) was ascertained, because I could get seeds on it by selfing. It has 
produced in 1919 the i'Vofispring containing 1 magenta, 28 reds, 12 oranges 
and 17 whites, thus proving itself to be heterozygous. We may consider the 
red examined to have been of the constitution CCUvbb, 1 and that it has 
changed into CcHrbb before the formation of gametes ; the male and the 
female gametes Cflb, elib, Crb, crb are formed, and their mating has 
given rise to 29 reds: 12 oranges: 17 whites, theoretically 32’6±:3'8 : 109 ± 
3*0:14-5 ±3 - 8 on 9:3:4 basis, if we will count 1 magenta as 1 red; 1 
magenta was formed because at least one gamete CRJB was produced by 
reversion, so that the whole process consists, exactly as in No. 29 (Table 
VIII, discussion p. 127), in the “ loss-mutation ” combined with reversional 
change. 
In the Table VI, A (p. 109) we have seen that the i^-offspring ex: 
white-II x orange No. 2 contains 81 magentas, 20 oranges, 34 whites and 1 
red, 136 in all. They are the total of all the offspring derived from .12 
-piants by selfing. Of these 12 families of the i^-offspring 11 contain no 
red at all, so that the one red under discussion belongs to one family 
composed of only 7 plants, viz. 5 magentas, 1 red and 1 orange. 2 The 
occurrence of 1 red in this family might perhaps be due to the partial linkage 
similar to that in the case just stated above, but no sure conclusion can be 
drawn here on account of the small number of individuals. 
Bucl-mutations have been observed, though not frequently. They are 
below simply described, but no discussion will be made, this being impossible 
1 Since the factors R and 15 were linked according to the series 5 :1:1: 5 or 6:1:1: 6 
in the preceding generation, we should have in the present case the reds of the composition 
CCRrbb and CCRRbb in the ratio 10:1 or 12 :1 respectively, and it is quite natural that 
we have now CCRvbb, because only two reds were examined. 
2 Whites are wanting in this small family, perhaps because all seeds which will give 
rise to whites failed to germinate. 
