238 
Fishery Bulletin 11 6(3-4) 
for fishing vessels. Such context and analyses can help 
fishery managers ensure that the Hawaii-based long- 
line fishery remains both ecologically and financially 
sustainable. 
Acknowledgments 
The authors are grateful for insight from E. Portner, C. 
Boggs, M. Donahue, A. Timmermann, and K. Edwards 
that greatly improved this manuscript. We thank the 
scientific observers of the Pacific Islands Region for 
their ongoing collection of longline fishery data. We 
also thank A. Tomita for help in formatting interna¬ 
tional fisheries data. This article is contribution 10356 
of the School of Ocean and Earth Science and Technol¬ 
ogy, University of Hawaii at Manoa. 
Literature cited 
Aires-da-Silva, A., and M. N. Maunder. 
2015. Status of bigeye tuna in the eastern Pacific Ocean 
in 2014 and outlook for the future. Inter-American 
Tropical Tuna Commission Stock Assess. Rep. 16, 13 p. 
lAvailable from website, accessed October 2016.] 
Bigelow, K., M. K. Musyl. F. Poisson, and P. Kleiber. 
2006. Pelagic longline gear depth and shoaling. Fish. 
Res. 77:173-183. 
Boggs, C. H. 
1992 Depth, capture time, and hooked longevity of 
longline-caught pelagic fish: timing bites of fish with 
chips. Fish. Bull. 90:642-658. 
Bopp, L., L. Resplandy, J. C. Orr, S. C. Doney, J. P. Dunne, M. 
Gehlen, P. Halloran, C. Heinze, T. Ilyina, R. Seferian, et al. 
2013. Multiple stressors of ocean ecosystems in the 21 st 
century: projections with CMIP5 models. Biogeoscienc¬ 
es 10:6225-6245. 
Cabre, A., I. Marinov, R. Bernardello, and D. Bianchi. 
2015. Oxygen minimum zones in the tropical Pacific 
across CMIP5 models: mean state differences and cli¬ 
mate change trends. Biogeosciences 12:5429-5454. 
Cox, S. P. T. E. Essington, J. F. Kitchell, S. J. D. Martell, C. 
J. Walters, C. Boggs, and I. Kaplan. 
2002. Reconstructing ecosystem dynamics in the central 
Pacific Ocean, 1952-1998. II. A preliminary assessment 
of the trophic impacts of fishing and effects on tuna dy¬ 
namics. Can. J. Fish. Aquat. Sci. 59:1736-1747. 
Federal Register. 
2016 Papahanaumokuakea Marine National Monument 
expansion. Fed. Reg. 81:60227-60234. GPO, Washing¬ 
ton, D.C. Available at website 
Garcia, H. E., R. A. Locarnini, T. P. Boyer, J. I. Antonov, A. V. 
Mishonov, O. K. Baranova, M. M. Zweng, J. R. Reagan, and 
D. R. Johnson. 
2013. World ocean atlas 2013. Volume 3: dissolved oxy¬ 
gen, apparent oxygen utilization, and oxygen satura¬ 
tion. NOAA Atlas NESDIS 75, 27 p. 
Gilman, E., M. Chaloupka, A. Read, P. Dalzell, J. Holetschek, 
and C. Curtice. 
2012. Hawaii longline tuna fishery temporal trends in 
standardized catch rates and length distributions and 
effects on pelagic and seamount ecosystems. Aquat. 
Conserv. 22:446-488. 
Harley, S., N. Davies, J. Hampton, and S. McKechnie. 
2014. Stock assessment of bigeye tuna in the western and 
central Pacific Ocean. West. Cent. Pac. Fish. Comm. 
WCPFC-SC10-2014/S A-WP-01, 115 p. [Available from 
website, accessed October 2016.] 
Howell, E. A., D. R. Hawn, and J. J. Polovina. 
2010. Spatiotemporal variability in bigeye tuna ( Thun- 
nus obesus) dive behavior in the central North Pacific 
Ocean. Prog. Oceanogr. 86:81-93. 
Kitchell, J. F., T. E. Essington, C. H. Boggs, D. E. Schindler, 
and C. J. Walters. 
2002. The role of sharks and longline fisheries in a pelagic 
ecosystem of the central Pacific. Ecosystems 5:202-216. 
Kume, S. 
1969. Ecological studies on bigeye tuna—V: A critical re¬ 
view on distribution, size composition and stock struc¬ 
ture of bigeye tuna in the North Pacific Ocean (north of 
16°N). Bull. Far Seas Fish. Res. Lab. 1:57-75. 
Lehodey, P, I. Senina, J. Sibert, L. Bopp, B. Calmettes, J. 
Hampton, and R. Murtugudde. 
2010. Preliminary forecasts of Pacific bigeye tuna popula¬ 
tion trends under the A2 IPCC scenario. Prog. Ocean¬ 
ogr. 86:302-315. 
Polovina, J. J., and P. A. Woodworth-Jefcoats. 
2013. Fishery-induced changes in the subtropical Pacific 
pelagic Ecosystem size structure: observations and theo¬ 
ry. PLoS ONE 8(4):e62341. 
Polovina, J. J., M. Abecassis, E. A. Howell, and P. Woodworth. 
2009. Increases in the relative abundance of mid-trophic 
level fishes concurrent with declines in apex predators 
in the subtropical North Pacific, 1996-2006. Fish. Bull. 
107:523-531. 
Saha, S., S. Nadiga, C. Thiaw, J. Wang, W. Wang, Q. Zhang, 
H. M. Van den Dool, H.-L. Pan, S. Moorthi, D. Behringer, 
et al. 
2006. The NCEP Climate Forecast System. J. Clim. 
19:3483-3517. 
Schaefer, K., D. Fuller, J. Hampton, S. Caillot, B. Leroy, and 
D. Itano. 
2015 Movements, dispersion, and mixing of bigeye tuna 
(Tliunnus obesus) tagged and released in the equatorial 
Central Pacific Ocean, with conventional and archival 
tags. Fish. Res. 161:336-355. 
Sibert, J., J. Hampton, P. Kleiber, and M. Maunder. 
2006. Biomass, size, and trophic status of top predators in 
the Pacific Ocean. Science 314:1773-1776. 
Stramma, L., G. C. Johnson, J. Sprintall, and V. Mohrholz. 
2008. Expanding oxygen-minimum zones in the tropical 
oceans. Science 320:655-658. Article 
Walsh, W. A., and J. Brodziak. 
2015. Billfish CPUE standardization in the Hawaii 
longline fishery: model selection and multimodel infer¬ 
ence. Fish. Res. 166:151-162. 
Walsh, W. A., P. Kleiber, and M. McCracken. 
2002. Comparison of logbook reports of incidental blue 
shark catch rates by Hawaii-based longline vessels to 
fishery observer data by application of a generalized ad¬ 
ditive model. Fish. Res. 58:79-94. 
Walsh, W. A., R. Y. Ito, K. E. Kawamoto, and M. McCracken. 
2005. Analysis of logbook accuracy for blue marlin (Mak- 
