228 Nichols. A roorphological study of Juniperus communis var. depressa. 
disappear after the reappearance of nucleoli in the resting nuclei. 
Later, after the homotypic division, small nucleolus-like bodies are 
frequently fonnd lining both surfaces of the cell plates. These, 
however, will be referred to presently* 
The body cell and the male cells to which it gives rise also 
present some interesting problems in connection with the nature 
of the so-called nucleoli, and it is evident from a study of these 
cells that the structures are of at least two sorts. The body cell 
possesses a large vacuolate nucleolus, which is probably homologous 
with the “plastin nucleolus” described by Coker (1903 b) in Taxo- 
dium. Düring the diyision of the nucleus, however, in contrast 
to the behavior of the nucleolus of the microspore mother cell, 
this body appears to shrink in size, although still maintaining its 
spherical shape, and is recognizable as late as the time when the 
chromosomes are becoming oriented at the equator (figs. 56, 57). 
Subsequent to this diyision prominent nucleoli of the same type 
reappear in the daughter nuclei. In addition to these structures 
there are present in the male cells small nucleolus-like bodies, the 
‘pseudpnucleoli’ of Noren (1907). These are distributed through- 
out the nucleus, and in view of the absence of any visible chro¬ 
matin are regarded by Lawson (1904, 1907) and others as chro¬ 
matin granules. Outside the nuclei of the male cells, immediately 
after their formation, and lining the cell plate on both sides there 
may be found in favorable material granulär or lumpy masses 
which likewise take the chromatin stains, and which are probably 
of the same nature as those referred to in the microspore mother 
cells. These masses are frequently very conspicuous in the young 
male cells (fig. 58) but usually disappear soon after the laying 
down of the dividing wall. They are doubtless homologous with 
the “plastin granules” described bv Coker (1903 b) in Taxodium, 
and it is doubtful whether they ought to be interpreted as nucleoli. 
In the egg nucleus the nucleolus-like structures are still more 
numerous and are exceedingly difficult to interpret. Frequently 
the entire chromatic content of the nucleus seems to have resolved 
itself into nucleoli and pseudonucleoli, yet even after the study of 
a large number of preparations one is unable to formulate any 
satisfactory conclusions as to the nature of these structures. That 
the pseudonucleoli are different from the typical nucleoli, however, 
seems obvious. Some at least of the former appear to be definitely 
associated with the reticulum, while in the mature egg nucleus 
they are the only parts which take the chromatic stains with 
avidity. It is not improbable that, as Wäger (1904) suggests, the 
pseudonucleoli “form a part of the nuclear network in which chro¬ 
matin or chromatin substance may be stored and possibly to some 
extent elaborated”. 
There is little doubt that many features in these nuclei which 
appear to be normal, and which have been described as such by 
various writers, are artefacts, and that the appearances seen in 
fixed and stained material differ greatly from the structures present 
in the living nucleus. Chamberlain (1906), after a study of the 
